Preface Plant protection is an integral part of crop husbandry without which the desired production can not be achieved. Production of food grain has tremendously increased in the last 50 years and it has touched the level of about 210 million tones presently, which is a matter of pride for Indian agriculture. We have not only enough food production but are also in a position to export it to other countries.     One can not deny the fact that heavy application of pesticides was made to save the crops from ravages of pests, diseases and weeds. But the indiscriminate use of pesticides has polluted the environment and affected our health adversely beside deleterious effects on non-target organisms. More and more and more insects are being reported resistant to different insecticides year after year posing a serious threat to agriculture. However, there has been work also on alternate control measures like biopesticides, varietal resistance, transgenic plants etc. for a better pest management strategy. The present book “Entomology: Novel Approaches” gives a vast knowledge about the progress made in India on different entomological aspects. Scientists from different disciplines of entomology will only be able to collaborate effectively if they have sufficient knowledge and understanding of newer technologies of pest management. This will only be possible if the necessary information is available in appropriate form. It has been our intension that this book should provide a source of information on the research techniques and practices of pest management, advanced genetic and biotechnological researches, new pest management technologies on different crops, pesticidal contamination status in environment. The aim of this book is to provide an overview of insect pest management, highlighting the recent developments, for teachers, students, researchers and extension workers engaged in pest management strategies. Views expressed and information provided in the present book are contributor’s opinion. We are thankful to all scientists and experts who have responded positively and have contributed the chapters.

Introduction The history of Integrated Pest Control goes back much farther than the mere coining of the term in the early fifties and the subsequent elaboration and clarification of the concept. The origins are deeply rooted in the evolution of pest control practices as developed by Entomologists and Plant Pathologists in the Nineteenth Century. Human history is a series of attempts to gain increasing control over the environment. At first this control was minimal to the degree that poor shelter and unstable food supplies imposed severe population constraints. The gradual gain in man’s capacity to control his environment parallels the gradual rise of civilization. But as man aggregated into villages near rivers and planted crops near by, he encountered increasingly severe attacks by pests on the crops. For thousands of years, man could do nothing about these pests but appeal to the power of magic and a variety of gods. For the most part, early humans had to live with and tolerate the ravages of plant diseases and insects, but gradually they learned how to improve their conditions through “trial and error” experiences. These improvements included the beginnings of pest control or pest management, the most preferred term today.

INTRODUCTION Management of several of the important insect-pests of crops is becoming increasingly difficult owing to a number of factors by which pests have been able to offset the effect of control interventions. Several of the present day pest management problems may be ascribed to the excessive and indiscriminate use of chemical pesticides during the last about three decades. Routine application of insecticides for controlling insect-pests, without taking into consideration whether economic damage to the crops is being inflicted by the pest or not, has resulted into several problems like pest resistance, pest resurgence, pest replacement, killing of natural enemies of pests, presence of toxic residues in food, and soil and environmental pollution, etc., apart from increase in the cost of plant protection. Entomologists have been trying to find solution to these problems through integrated pest management of which monitoring of insect activity through regular surveys is an important component. In fact, knowledge of insect activity is indispensable since pest management system can not operate without accurate estimates of pests and natural enemies population or without reliable assessment of plant damage and its effect on yield.

Introduction Humans have inhabited the earth for nearly 2 million years. For over 90 per cent of this period, the maximum population numbered about 200,000 and the growth rate was only about 0.01 percent per year. The use of fire and the development of tools were the first major changes affecting human population growth which increased to about 5 million individuals about 100,000 years ago. The population remained fluctuating about this mean value for the next 90,000 years depending upon the food supply. Up to this time people were hunterer-gatherers and depended exclusively on the natural ecosystem for their food. Then about 10,000 years ago, humans began to cultivate crops for food. With established agriculture came an increased and stable food supply. This contributed to an increased growth rate of the human population (Dhaliwal et al., 1998).

INTRODUCTION The first significant economic contribution of host plant resistance in agriculture was made in 1890 when European grape vines were successfully grafted on resistant root stock to save the French Wine industry from viteus vilifaliac (Fitch). Studies from the past century have led to the development of resistant varieties in about 15 crops to about 50 key insect pests. During the last 20 years alone, more than 500 insect resistant cultivars of rice, maize, cotton, sorghum, alfalfa and wheat have been developed and released world wide (Dhaliwal and Arora, 2001). The insect resistant varieties are grown on millions of ha. annually and help the farmers to save billions of dollars in insecticide costs (Dhaliwal and Arora, 2001). The role of Host Plant Resistance in Integrated Pest Management is discussed below.

Introduction Advent of agriculture in India dates back to around 2000 BC, the period of Indus Valley Civilization. South and East Asia also witnessed the start of agriculture around the same time. The crops were grown only during the monsoons. Winters served as a closed period for crops as well as for the pests. Even at that time a number of insect pests were known to cause serious damage to different crops. Agriculture has been revolutionized during the few decades since World War II (Kupatt et al., 2005). Introduction of high yielding varieties and associated technology resulted in tremendous increase in the production from the same piece of land. The results of these changes include a dramatic reduction in time and resources needed to provide an unprecedented quantity of quality food and fiber. The initial introduction of plant protection agents as wonder material did not lasted long, as very soon their ill effects on human health and environment were witnessed.

INTRODUCTION   In India tremendous progress has been made in agriculture. In absence of check on population growth, it may cross 1.5 billions by 2030. As a consequence per capita availability of land 0.13 ha will further reduce to 0.1 hectare in 2020 A.D. The demand of growing population for food and fibre require country to produce higher yields from presently cultivated land. Currently an average of 2790 calories of food available each day for every human on the planet, (23 per cent more than in 1961 and enough to feed every one). The production targets and required yield to feed the ever increasing current population with present available resources will require the growth rate of 2.35 and 2.2 per cent in rice and wheat, respectively and as high as 4.45 and 3.38 per cent for pulses and oil seeds, respectively. This requires national average yield to be increased by 30-50 per cent, which is not a cakewalk. India will require producing additional 5-6 million tones annually to keep pace with ever-increasing national food and nutritional requirements. Recurrent famines, starvation death and lack of resources led to green revolution, which as of now has turned our begging bowl to agriculturally advanced cereal exporting country.

INTRODUCTION While the world has been changing during the last few years both politically and economically in unexpected and remarkable way, food security remains an unfulfilled dream of billions of people. With the world population crossing six billion marks, the demand for food grains is increasing at an alarming rate. Increasing population, on the other hand, has led to decrease in cropping area. In order to meet the global food demand we have to increase the productivity. Insects have always been a major cause of yield-loss in agriculture. They account for 20-30% of crop-loss. Several approaches e.g. physical, chemical and biological have been used with varying degree of success to reduce such losses.   The insect pest management is one of the most important yield destabilizing factors in almost all the crops. While for some insects, genetic sources of resistance are available in the respective plant germplasm, for many others germplasm is almost devoid of effective resistance genes. It is because of this factor that conventional breeding strategies have not been successful in breeding insect pest’s resistant varieties. The only means available at present for managing the insect is extensive application of insecticides and pesticides. Excessive usage of the pesticides causes, on one hand, environmental pollution and on the other, it forces the insects to develop resistance against these chemicals. Application of the pesticides also imposes additional economic burden on the farmers.

Introduction Pesticide in broad term refers to “killer of pests” but is an agent used to control, prevent, destroy, repel and mitigate a pest. Pesticides are thus poisonous substances intentionally applied in situations, which harbour so-called pests. Agriculture including poultry and veterinary (i.e. raising of crops and protection of produce and storage), health (public places-hospitals, restaurants, gardens cinema halls etc.), transportation (shipment, air rail and road transport) are some areas where pesticides are used extensively as spot application, fumigation, aerial and field sprays, soil treatments etc. Plant protection comes to be recognized as an equally important factor of agriculture production as improved seeds, fertilizers and irrigation. Pesticides over time get accumulated in the environment and contaminate all systems i.e. air, water, soil, plants, animals and micro-organisms on being transported from one system to another leading to biomagnifications. The fate of a pesticide in the environment and the entities, which come in contact, is depicted in the Fig. 1 reflecting the pollution and contamination. Pesticides during manufacturing, transportation, storage and actual use enter the biotic and abiotic components of the environment through air, water and soil and disturb the cohesion causing great disaster sometimes. Pollution is defined as undesirable change in physical, chemical and biological characteristics of air, land and water. This change may or will harmfully affect human life or that of desirable species, industrial process, living conditions and cultural assets. It may or will waste or deteriorate our raw material resources. Pollution increases burden of human society causing loss of resources through unnecessary wasteful exploitation, the cost of pollution abatement and deterioration of human health.

Introduction The world of living organisms essentially consists of the kingdoms of plants and animals. Myriads of organisms representing these two, interact among them, ranging from symbiosis and mutualism to competition and predator-prey interactions. The last mentioned process is one the important components of ecological dynamics, that determine the balance of nature or equilibrium among the biota. To this category belong the predators, parasites (parasitoids), pathogens, and so on, all of which live at the cost of another organism, without affording any benefit to the ‘victim’, hence the victim is termed the ‘prey’. In an agroecosystem, when an organism, explodes into an unusually high number, devouring cultivated plants it often becomes a ‘pest’ from a man’s point of view. Sometimes the pest can be a human nuisance too. It is here, that the role of predators and parasitoids become relevant as biological agents against the pests.   A lion, which preys on herb-feeding antelopes, is also a predator. But, its roles become restricted to maintaining the balance of the antelope population. Left undeterred, antelopes can explode to levels that can threaten grasslands. Here lions play a very crucial role in maintaining (through hunting antelopes) the equilibrium of the grasslands, just sufficient for these plants to surge with every rain. Otherwise, the grasslands potentially may get denuded by the antelopes, or conversely it may override other ecosystems. This is part of the ‘food web’ and is found in every ecosystems; desert, aquatic, mountains, forests, agro-ecosystems, etc.

INTRODUCTION “An insect is like an old-fashion soldier: it does what it is ordered to do, and when it has no orders it does nothing” (a citation from Wright, 1963). The orders are very often given as chemical signals that induce certain behaviour or change the course of development in the insect. Chemical communication appears to be the primary mode of information transfer in insects. These chemicals are able to modify the behaviour of a perceiving organism at sub-micro/nanogram levels and are known as semiochemicals. The term is derived from the Greek Word Simeone, meaning a mark of a signal. These behaviour modifying chemicals are divided into two broad categories.

Introduction Although the concept of IPM, with emphasis on minimum use of pesticides; is well established and widely practiced in India pest population still continue to be unabated, often reaching to alarming levels. Out breaks of Spodoptera litura and hairy caterpillar, Amsacta moorei have been recorded in soybean and kharif crops recently during 2002 & 2004 in Madhya Pradesh and Rajasthan, respectively. Outbreaks of Helicoverpa armigera populations are well known, appearing more frequently since past few years (1987-88, 1995-96, 1997-98 & 2001) doing extensive damage in cotton and pulses. in many parts of the country. Failures in controlling these populations, even by most potent insecticides, have raised a serious doubt on the success of present day pest management practices aimed to kill the pest in immature larval / egg stage, when infestation spreads out in large areas. Insecticidal treatments require wide coverage of crop area to control them effectively. Insecticide resistance in pest species and destruction of natural enemies are obvious problems, since chemicals are applied indiscriminately to entire crop ecosystem including the non-target beneficial bicontrol species. How to minimize use of insecticide in pest control is often a question of major concern to the entomologists.

INTRODUCTION Insecticide resistance poses a severe threat to agricultural productivity in India and in other countries of the world. Humans have had the ingenuity to develop thousands of pesticides to protect foodstuffs, livestock, and health and the use of these poisons has triggered a rapid evolutionary response in many target species and some nontarget species of invertebrates, microorganisms, higher plants and vertebrates. Insecticide exposure led to nullify their toxic effects through genetic selection for individuals with the biochemistry or behaviour. Continual development of new compounds and its greater use and increase in input insecticide cost is mainly due increase in resistance. Reduction in insecticide efficacy from insect resistance has major economic, environmental and human health implications. Integrated Pest Management (IPM) is a convenient descriptor for wide range of still-evaluating pest control program that emerged from chemical prophylaxis practiced. IPM theory has fostered a transition from primarily chemical control to programs tempered with biological information and input tailored to each agro ecosystem, crop and pest situation in production.

Introduction The introduction of high yielding varieties and hybrids in late sixties led to accumulation of insect pest problems in cereals. The varieties and hybrids developed during green revolution era with the sole objective to enhance production to feed increasing population were better in quality parameters than traditional locals but were highly susceptible to insect pests. Over 200 pests attack different cereals and insects that were not known or were of minor importance have now become major pests and several new insect pests have appeared on cereals. Out breaks are frequent due to several obvious reasons. Major pests are serious, perennially occurring and persistent species which cause considerable loss in yields every year in large areas require control measures. Occasional pests attack the crops sporadically but cause potential damage and sufficient loss, though in some localities or only in some seasons, so as to warrant human intervention.

Introduction Rice is the staple food of about 50% of the world’s population that resides in Asia, where 90 % of the world’s rice is grown and consumed. In the International year of Rice celebrated in 2004 the theme was “Rice is life” which reflects the importance of rice as a primary food source of human beings. In the world, annually rice is cultivated in about 151.54 million hectares with an annual production of 593 metric tonnes and average productivity of 3.91 tonnes/ha (Rai, 2004). From its Asian homeland, it is now cultivated in 113 countries and plays a variety of roles related to important aspects of food security as well as rural and economic development. In Asia this crop is grown in 136.07 million ha followed by 7.67 million ha in Africa and 5.09 million ha in Latin America with a production of 539.84, 16.97 and 19.54 metric tonnes and productivity of 2.97, 2.21 and 2.84 tonnes, respectively (Rai, 2004). In India, rice is the staple food for more than 65 % people (Pasalu and Katti, 2004). Rice is grown under various geographical, climatic and cultural conditions. The current level of rice production in the country is around 90 million tonnes. Based on the present rate of population growth @ 1.5 % and per capita consumption of 250 g of rice/day in the country, the demand for rice production is expected to be about 100-105 million tonnes by 2010 and 130-140 million tonnes by 2025. To achieve this target, it requires increasing productivity per unit area by the adoption of modern and intensive agricultural practices, such as use of high yielding varieties and improved cultural and management practices. Experience of post green revolution period in last four decades indicates that these practices often aggravate other biotic constraints such as insects, diseases and weeds. However, insect pests are considered to be the major limiting factor in enhancing the productivity of rice.

INTRODUCTION Cotton is an important commercial crop and plays a key role in the economy of country. Cotton is grown in different states of India viz Punjab, Haryana, Rajasthan, Gujrat, Maharashtra, Madhya Pradesh, Andhra Pradesh, Karnataka and Tamil Nadu. In India the area under cotton crop is 8.76 m ha and the productivity is 436 Kg lint/ha as against the world average of 682 kg lint/ha (Anonymous, 2005). The productivity of cotton is variable in the country in the range of 287 to 720 kg lint/ha (Anonymous, 2005). The major constraints in cotton cultivation in India are as follows   MAJOR CONSTRAINTS IN COTTON CULTIVATION IN INDIA     i.    Abnormal weather conditions      ii.    Cultivation of cotton under rainfed condition      iii.    Cultivation of large number of susceptible and undescript cultivars with varied growth pattern.      iv.    Poor soil health     v.    Long duration of crop      vi.    Long sowing period      vii.    Indeterminate nature of cotton plant      viii.    Intensive cultivation      ix.    Imbalance use of fertilizers      x.    Non judicious use of irrigation water      xi.    Higher incidence of insect pests due to 

INTORDUCTION Spices are strongly flavored aromatic substances of plant origin which are commonly used for seasoning, cooking food and for preserving other food stuffs. There are about 70 species of spices grown in various parts of the world. Those which are used most commonly throughout the world are grown in India and other parts of South Asia. The seed spices are annual herbs, whose dried fruits or seeds are mostly used as spices. The seed spices are used for culinary, confectionary, perfumery, cosmetics and pharmaceuticals industry. There are about 20 seed spices grown in India (Table 1) and nine out of them are prominent. The most important among them are belongs to family Apiaceae (coriander, cumin, fennel etc). about 9 lakh ha area is growing in seed spices in the country with annual production of about 5-6 lakh tones (Anonymous, 2003). India is the world’s largest producer, consumer and exporter of seed spices. The foreign exchange earned through export of seed spices is more than Rs 200 crores annually. India exports raw spices as well as value added items to nearly 70 countries in the world and is meeting 51% of the global demand of these commodities. There is good potential for increasing export of seed spices, if production as well as quality is increased. The level of productivity in various seed spices is comparatively low as compared to other countries. There are many production constrains attributed to low productivity. The lake of suitable high yielding varieties suiting to different agro-ecological regions and more incidences of diseases and insect pests. Among different insect pests, coriander aphid, Hydaphis coriandari (Das) is one of the most destructive and widely distributed pest of coriander (Jain, 1984 and Jain and Yadav, 1989). Recently brown wheat mite was observed attacking coriander and other spices (Jain and Yadav, 1986 and 1989).

Introduction In India, climatic conditions are very diverse and fruit crops of temperate, sub-tropical and tropical regions are grown. Frequently, the fruit crops of one regional environmental requirement are grown in another region. This type of crop husbandry has some typical pest problems as the insect and mite pests have easy access to any of these regions depending upon the fruit crop. The application of management approaches in isolation of insect pests on fruit crops demand a unified strategy to suppress the different pest populations under the concept of ‘crop management’. Thus, to sustain the crop management for the better fruit quality, a judicious integration of safe chemical/biopesticide molecules must be used with other non-chemical approaches namely host plant resistance, bioagents, cultural and mechanical practices etc.

Introduction India ranks second in the production of vegetables after China. The existing area under vegetable cultivation in India is around 4.5 million ha. There is a need of around 5-6 million tons of food to feed 1.3 billion India’s population by 2020 AD. The major constraint in vegetable production, include the extensive crop losses due to increased pest infestations directly or due to viral diseases vectored by insects. The extent of crop losses in vegetables varies with the plant type, location, damage potential of the pest involved and cropping season. Vegetables are more prone to insect pests and diseases mainly due to their succulence as compared to other crops. The crop losses to the tune of 40 per cent have been observed in vegetable crops. The increased demand and limited productivity of local cultivars, led to introduction of high yielding varieties and hybrids. As a result, dramatic changes in pest scenario occurred like minor pests assuming major status (eg., Serpentine leaf miner, Liriomyza trifolii a serious problem in tomato hybrids). Chemical pesticides are the sole panacea to mange the pest by vegetable farmers even today. Indiscriminate use of pesticides has led to severe ecological consequences like destruction of natural enemy fauna, effect on non-target organisms, pesticide residues in vegetables and resistance of the pest to pesticides. The modern pest management is aimed at educating the farmers on various issues of pest management like identification of the pests, natural enemies, judicious use of pesticides with alternate methods of pest control like cultural, mechanical, biological and behavioral (pheromones) management.

INTRODUCTION Potato is a crop of major significance in human nutrition, ranking fourth in world production, after wheat, maize and rice. Potatoes are grown on over 44 million acres in more than 125 countries, with annual production of about 250 million tons. Potatoes are grown across India, with production concentrated in several areas of the North Indian plains and West Bengal. Indian potato production is significant and accounts for 8.5 per cent of the world total (Dahiya et al., 1999). The potato has hundreds of recognized pests including many insects, mites, nematodes and rodents. Some of these were transported to new locations alongwith seed tubers. Others were already present in locations where the potato was introduced and then proliferated on this new host plant. Because a potato crop is vegetatively propagated from tubers, which easily carry some pathogens and pests, many pest problems have followed potatoes to areas wherever they are grown. The potato tuber moth was introduced accidentally into India from Italy in 1906, probably through seed potatoes (Lefroy, 1907), and by the middle of century had become well established throughout India (Kumar and Nirula, 1967). Of all the major food crops, potato is heaviest user of chemical pesticides, which often total up to 20 per cent of its cost of production (Anonymous, 1991).

Introduction Rapeseed-mustard is attacked by more than 43 insect species. Out of which, mustard aphid, Lipaphis erysimi (Kaltenbach); mustard sawfly, Athalia proxima (Klug); painted bug, Bagrada hilaris (Burnmerister); pea leaf miner, Chromatomyia horticola (Goureau) and Bihar hairy caterpillar, Spilarctia obliqua (Walker) are of major importance. Other pests are of sporadic occurrence. Status of important insect pests attacking rapseed mustard in India have been given in detail by Bakhetia and Arora (1993) and Singh 2005. The occurrence of various insect pests on rapeseed-mustard is stage-specific. This has been a gift provided by nature to help avoid inter-competition among them for their fullest exploitation of the host. The insect pests attacking rapeseed- mustard as given in Table 1 and at various stages of the crop in Table 2.

INTRoDUCTION In developing countries, cereals and legumes are the principal source of daily energy and protein intake. The major economic loss caused by grain infesting insects in not always the actual material they consume, but also the amount contaminated by them and their excreta which makes food unfit for human consumption. Today million of tones of the food grains are damaged by the ravages of the pests that not only cause quantitative but also qualitative changes in food grains. Stored product insects depress the food value of a product, introduce pathogens, parasites or toxins or make it less palatable. Insect infestation in high in the tropics and may lead 100 per cent damage in severe cases.   In order to cope with the requirement of the increasing population and food problem, at present as well as for the decades to come, the world will have to raise agricultural production very substantially, both by increasing the production and by reducing losses on account of post harvest spoilage especially of cereals and legumes which are the primary sources of food for men kind. This is a major problems during agricultural production, food transfer and processing. Million tonnes of grain worth several hundred corers of rupees every years are either damaged or loss for want of knowledge of scientific methods of saving the foodgrains during storage. In India, post-harvest losses caused by the unscientific storage, rodents, insects, micro-organisms, moisture etc. account for about 10 per cent of total foodgrains (Anonymous, 1971). A world survey by F.A.O. indicated about 5 per cent loss of cereals, in storage annually (Herford, 1952).

Introduction Indiscriminate use of acaricides not only causes the development of resistance in phytophagous mites but are also harmful to their natural enemies, therefore, poses a serious problem of pest resurgence. Hence, worldwide attention has been focused to utilize the natural biocontrol agents and minimize the use of chemical pesticides for the control of mite pests in agriculture with a view to develop a suitable IPM strategy. Among natural enemies of phytophagous mites, predatory mites have been found to be most effective and efficient tool in IPM programme because of the fact that many of these are not only abundantly available in nature but are also voracious feeders and have proven ability to suppress the pest mite population below economic injury level. Beside this, predatory mite can locate, run and catch hold of the pest mites even the pest mite is hiding in rolled leaves, under webs or even in the galls or erinium, etc. Some of the predatory mites require very low population of pest mites as prey, for their survival and multiplication, which is an added advantage for potential predation. Some predatory mites such as Typhlodromus pyri have good synchronization with the prey mite, Panonychus ulmi as, both hibernate and remain active in orchards.

INTRODUCTION Phytophagous mites have long been considered as potentially serious pests of a wide range of food and fibre crops, and ornamentals. They are covered under subclass Acari of class Arachnida and phylum Arthropoda. Mites unlike insects are 8-legged creatures and lack compound eyes, antennae and wings. Plant feeding mites measure from 200µ to 600µ. Majority of the plant feeding mites belong to suborder Prostigmata, and phytophagous mites belong to family Tetranychidae (spider mites); Eriophyidae (bud or gall mites) ; Tenuipalpidae (false spider mites); Tarsonemidae (broad mites) and Tuckerellidae, in the decreasing order of their importance. Before one can develop integrated management of phytophagous mites in different agri-horticultural ecosystems, knowledge about the following fundamentals of phytophagous mite management principles is a big must.

Introduction Leafhoppers belong to the family Cicadellidae and constitute one of the largest families in the exopterygotan Hexapoda. Recent estimates of species for the world range between 35,000-45,000 species though little over 22,600 species are now described (Dietrich, 2004a). The leafhoppers belong to the order Hemiptera and suborder Auchenorrhyncha.   Leafhoppers can readily be recognized from other members of the Auchenorrhyncha by the presence of two or more rows of spines on hind tibiae and by the pronotum not extending back over abdomen. Their close relative, the Membracidae, have pronotum extending back over the abdomen. From planthoppers (Delphacidae) with which they are often confused, they can be recognized by the position and structure of antennae. The leafhopper antennae are placed between eyes compared to beneath the eyes in planthoppers; the second segment of the antenna (pedicel) is small without externally visible sensillae in leafhoppers compared to enlarged pedicel with prominent sensillae in the planthoppers. The movable large spur at the apex of hind tibia in plant hoppers is absent in the leafhoppers.

Introduction India is the largest developing economy of the world and about 70 per cent of its population is dependent directly or indirectly on this profession.  To meet the food requirement of the ever-multiplying population of this country, agriculture production has to be maximized by using genetically improved varieties, fertilizers improved implements, seed, and other agronomical technologies. Beside these improvements the agriculture production has remained static for the last decade. For further improvements in agricultural production we have to adopt comprehensive strategies. One of such strategies is the pollination management that is a very vital process of agriculture crop production. Pollination is a process involving reproductive parts of a flower and an external agent in the form of a vector. According to Prof. E.C. Martin, the value of bee pollination of fruits, vegetables and seeds in the United States of America totaled very close to $8 billion in the 1960s; the figure 20 years later is probably close to $20 billion.

Introduction Many insets inhabit the soil and participate in different activities which will result in changes in soil physico-chemical properties. Thus soil is a very good habitat for some insects. Several insects spend either complete or few stages of their life cycle in soil. Soil provides protection from predators for many soil dwelling insects especially in immature stages. Few insects also undergo diapauses in soil. Soil insects include phytophagous, scavengers, predators and parasites. Among these, soil macro fauna especially termites and white grubs are very important pests of many crops and their activity also brings many changes in soil ecosystem. Termites Termites (Order: Isoptera) are eusocial, polymorphic and live in small to large colonies. The colony consists of different castes viz., functional reproductive (queen and king), sterile workers and soldiers, which are morphologically distinct.  The tasks of the latter two castes include foraging, nest building and care of eggs and young ones and defense. All species maintain symbiotic relationships with microorganisms, which aid in digestion. Few species build the often-spectacular mounds or arboreal nests and others build subterranean nests. They constitute important group of soil fauna and play a major role in the arid and semiarid tropics. Worker castes are dominant and are responsible for major functions in the colony. During the process of food collection, the workers generally attack the plants that are weakened due to some internal growth factors or environmental causes such as drought or poor soil conditions (Roonwal, 1978).

Author Index Agarwal, V.K., 453 Bajpai, n.k., 181 Batra, R.C., 333 Bhargava, M.C., 133, 319, 425 Bhullar Manmeet B., 461 Chakravarthy, A. K., 97 Chandel, R.S., 377 Chandla, V.K., 377 Choudhary, R.K., 133,  319, 425 Dhiman, K.R., 377 Dhooria, M.S., 461 Dubey, O.P., 121 Gupta, Rakesh, 109 Jaglan, R.S., 1 Jain, P.C., 133, 319, 425 Jana, R.S., 499 Jena, b.C., 267 Kaushik, H.D., 499 Kavadia, V.S., 145 Kishore, Prem, 245 kumar, Ashok, 181 Kumar, N.G., 509 Kumar, Vinod, 377 Misra, H.P., 267 Nagaraju, D.K., 165 Nath, Paras, 43 Nimbalkar, S.A., 287 Pathrose, Berin, 211 Purwar, J.P., 399 Rajagopal, D., 97 Sachan, G.C., 399 Saini, R.K., 19 Satpathy, S., 353 Sharma, Ajay, 109 Sharma, Ashok, 453 Sharma, D.R., 333 Sharma, O.P., 121 Sharma, R.K., 245 Shivalingaswamy, T.M., 353 Singh, Ram, 1 Srivastava, Chitra, 211 Vaishampayan, S.M., 193 Verghese, Abraham, 165 Viraktamath, C.A., 477 Yadav, P.R., 19