eChapters

Breeding objectives genetic resources and techniques for breeding

• There are countries that consume 100 to 150 kg yearly per capita while others which do not even make it to 10kg.
• Genetic variability is the raw material of the plant breeder.
• International network for the improvement of Banana and Plantain (1 NIBAP), Prance to conserve the available germplasm.
• Only pathogen gene-free planting material can be used for construction or distribution from the gene bank for further multiplication or utilization in any breeding program.
• Fruit breeders have therefore tended to utilize single gene variants in performance to polygenic variation.
• In general, for most commercial requirements, homozygous is inadequate by six or seven generations of selling (56 or 37).
• In the dominance hypothesis, it is assumed that the deleterious gene is normally recessive.
• The simplest approach for the selection of our breeders is mass selection.

Indian jujube (Ber) belongs to the genus Zizyphus and family Rhamnaceae having more than 600 species. The family includes trees with spine, shrubs and climbers. The original home of Ber is Myanmar and India. Many workers have tried to classify it on the basis of leaf, flowers and fruits characteristics. Some tried to divide into two groups i.e. having smooth leaves on both surfaces and having leaves downy beneath. Ber as a tree has been described it as small, much branched or a large shrub with dense spreading habit, bark dark gray, branches elongated, leaves with glabrous shining upper surface densely covered beneath with whitish hairy growth.

Ber has attracted sustained breeding attention owing to its wide adaptability, good quality and nutritive value. However, problems like poor shelf life, incidence of powdery mildew disease, fruit fly and fruit borer have hindered the extension of the area under this crop. In Chinese jujube, the problem of fruit cracking and ‘witches broom’ disease are the major hindrances in its cultivation. There are two major cultivated species viz., Ziziphus mauritiana and Ziziphus jujuba, which are widely distributed covering vast regions in Asia. The wild population found is heterozygous and extremely variable. As a result, many local selections were made, some of which have become widely recognized as cultivars. The success of any breeding programme depends on the extent of genetic variability in the source population. The assessment of variability is therefore a basic requirement of breeding programme. Since most of the plant characters of economic importance are governed by the group of genes and are highly influenced by environmental variation, it is difficult to judge whether the observed variability is heritable or due to environment. This necessitates optimizing of phenotypic variation into its heritable and non heritable components, so that the selection of parents

5.1. Characteristics of asexually propagated crops

Majority of them are perennials

Many of them show reduced flowering and seed set.

Invariably cross pollinated.

Mostly polyploids.

Many species are inter specific hybrids.

4.1. Synthetic varieties

Produced by crossing in all combinations, a number of lines that combine well with each other.

The genetic organization of cross pollinated crops are different from that of self pollinated crops, because of differences in the reproductive system. Open pollinated populations are heterozygous with free inter mating individuals, heterozygous at most of the loci. Heterozygosity is an essential feature of commercial varieties of out breeders. It is maintained during breeding or is restored at the end. Population improvement involves selection, generation after generation, with recombination of selected individuals. Its aim to increase the frequency of favorable alleles in the population. The improved forms may be used for commercial cultivation or as a base germplasm for the derivation of inbred lines. The list of cross pollinated plants is presented in Table 10.1. The factors that promote the cross pollination are dicliny, dichogamy, herkogamy, self-incompatibility and male sterility. The breeding methods which are commonly used in cross-pollinated crops are population improvement, hybrid and synthetic varieties.

Population Improvement

In cross pollinated crops heterozygosity must be restored in the end product of any breeding programme. It is classified into the following types:

3.1. Selection

To isolate desirable plant types from a population.

3.1.1 Progeny test

Evaluation of growth of plants on the basis of performance of their progenies

Real value of plant could be known by studying the progeny.

The prime object of plant breeder is to develop high yielding variety with superior characteristics. Selection is based on the phenotypes of the plants. Consequently the effectiveness of selection primarily depends on the degree to which the phenotype of plants reflects their genotype. These are all self fertilizing species. In these species development of seed take place by self fertilization. Hence self pollinated species are also known as autogamous species. Various plant characters such as homogamy, cleistogamy, chasmogamy, bisexuality etc. promotes self pollination. The self pollinated crops with their description are given in Table 9.1. The breeding methods used in self and often-self pollinated crops are discussed below:

Plant Introduction

Plant introduction consists of taking a genotype or a group of genotypes of plants into a new area or region or environment, where they have not being grown earlier. It is very simplest and fastest method of developing new variety. It is of two types:

Reproduction that does not involve the union of male and female gametes is known as asexual reproduction. Reproduction in which sexual organs or related structures take part but fertilization does not occur, or crops which are propagated asexually or by vegetative means are known as asexually propagated crops. Most of the fruit plants are propagated asexually which consist of large number of clones. Example: sugarcane, potato, sweet potato, banana, mango, citrus, pears, peaches, litchi, crops that propagates by asexual means. The procedure of selection used for asexually propagated crops is known as clonal selection, since the selected plants are used to produce new clones or superior clones can be isolated from local variety, introduced variety or inter crossed population.

Important breeding methods can be classified as follows:
    1.    Introduction
    2.    Selection
    a.    Mass selection: Done in self pollinated crops, cross pollinated crops and dioecious crops.
    b.    Individual plant selection: Done in self pollinated crops, cross pollinated crops (line breeding) without control of pollination, artificially self pollinated lines, dioecious plants and clonally propagated lines.
    3.    Hybridization
    a.    Crossing in self pollinated crops.
    b.    Single crosses, double crosses and synthesized varieties.
    c.    Varietal crosses.

Introduction
In recent years, there has been a growing interest in alternative nutritious food sources (McClements, 2020) and such group of foods that has gained significant attention is millets. Millets are a diverse group of small-seeded grasses that have been cultivated and consumed for thousands of years. They are rich in essential nutrients such as proteins, fibers, vitamins, and minerals, making them an important addition to a balanced diet. Millets are not only rich in essential nutrients but also have several health benefits. They are known to have nutraceutical properties, low glycemic index, making them suitable for individuals with diabetes. Additionally, millets are a good source of antioxidants, which can help in reducing the risk of chronic diseases such as heart disease and cancer. The diverse range of millet varieties also provides flexibility in incorporating them into various dishes, making them suitable for different culinary preferences and dietary requirements. Therefore, the usage of millets can play a crucial role in promoting sustainable and healthy eating habits.
Malnutrition continues to be a pressing issue worldwide, particularly in developing countries (Ashok et al., 2020). The lack of access to nutritious food is one of the main causes of malnutrition, leading to various health problems such as obesity, diabetes, cardiovascular diseases, cancer, and celiac disease (Birania et al., 2020). To address this issue, millets have emerged as a promising solution. Despite being nutritionally superior to other cereals, millets have traditionally been limited to consumption by the poor and those with conventional dietary practices. However, there is a growing recognition of the potential of millets in combating malnutrition and improving public health. Millets, such as finger millet, pearl millet, and sorghum, are often referred to as “superfoods” due to their high nutritional content (Singh et al., 2020). These grains are rich in protein, carbohydrates, fats, vitamins, minerals, and dietary fiber, making them an ideal source of essential nutrients (Ashok et al., 2020). Additionally, millets are easily available and costeffective, making them a viable option for addressing malnutrition on a larger scale (Birania et al., 2020). Furthermore, millets are resilient crops that can withstand environmental changes and water shortages, making them an ideal choice for sustainable agriculture. In this chapter, we delve deeper into specific aspects of millet nutrition, examining their micronutrient profiles, health benefits, culinary versatility, and potential applications in combating malnutrition and promoting sustainable agriculture. By understanding the nutritional richness of millets, we can uncover innovative solutions for improving human health and fostering agricultural resilience.

4.1. Flower and flowering in pigeonpea

Growth habit: Primarily, there are two major growth types in pigeonpea. These are determinate and non-determinate. The determinate growth in the pigeonpea plants is characterized by flowers in bunches at the top of the canopy (Fig 17a). In this case the epical growing points are transformed in to reproductive buds and bunches of flowers appear on the top of the canopy. Such cultivars are not popular as the management of insects such as Helicoverpa and Maruca becomes very difficult. Most of the cultivated types belong to another growth category called as non-determinate. In such genotypes the apical growing points remain vegetative and the plants keep growing under favorable conditions and it results in tall canopy with many spreading or compact branches. The inflorescence in this case emerges from the axil of secondary and tertiary branches (Fig 17b); and hang sideways. These types show less susceptibility to insects due to non-clustering nature of flowers.

17.1 Introduction

Almond or badam is a high energy nut. Almond kernels contain lipids (52.2%), protein (20%), carbohydrate (20.4%) and water-soluble sugars (4.4%). Irrigation affects almond yield and quality. Almond quality is primarily genotype dependent. Maturity at harvest can also influence kernel quality.

Aonla has been regarded as a sacred tree in India. The tree is worshipped as mother earth and is believed to nurture humankind because the fruits are very nourishing. It is also known as Amrit phal. Aonla is mainly recognized due to its high nutritive, medicinal and therapeutic properties. It contains high amount of vitamin C (500-1500 mg/100 g), it is also rich in pectin, iron, calcium and phosphorous. Dried fruits of aonla can be is used in curing hemorrhage, diarrhea, chronic dysentery, diabetes, jaundice, dyspepsia, cough etc. Recently, aonla has gained more popularity in arid and semi-arid region of the country because of its hardy nature, prolific bearing capacity and ability to grow under various adversities as a remunerative crop (Shukla et al.2002). It may be an important fruit in future due to its high productivity, suitability in wasteland particularly salt affected soil and sandy soil of arid ecosystem. Aonla is the main ingredient of Chavanprash and Triphala powder which is used for curing different abnormalities. The fruit contains a chemical substance gallic acid and leucoanthocyanin which have antioxidant property.

12.1 Introduction

Aonla (also called Amla, Indian gooseberry) is a tropical and sub-tropical fruit. It is mainly recognized due to its high nutritive, medicinal and therapeutic properties. It contains high amount of vitamin C (500-1500 mg/100 g); it is also rich in pectin, iron, calcium and phosphorus. Dried fruits of aonla are used in curing haemorrhage, diarrhoea, chronic dysentery, diabetes, jaundice, dyspepsia, cough etc. Aonla is the main ingredient of Chavanprash and Triphala powder. The fruit contains gallic acid and leucoanthocyanin which have antioxidant property. Its cultivation is more common and widely grown in Uttar Pradesh, Gujarat, Rajasthan, Madhya Pradesh and Tamil Nadu.

14.1 Introduction

Apple is the most widely grown temperate fruit of the world. It is considered as King of temperate fruits. It is rich in carbohydrate, calcium, phosphorous, iron, potassium, vitamin B6 . Apple is rich in antioxidant, flavonoids and dietary fibre. It is recommended to reduce the incidence of dental caries. The phytonutrients and antioxidants may help reduce the risk of developing cancers, hypertensions, diabetes and heart disease. Apple is mainly used for table purpose. The various processed products are prepared from fruit, e.g., juice, jelly, preserve, canned slice, wine, concentrated juice, cider and powder. It is widely cultivated in Afghanistan, Armenia, Azerbaijan, Bhutan, China, India, Indonesia, Iraq, Iran, Israel, Japan, Jordan etc.

20.1 Introduction

Arecanut (or betel palm) is chewed as narcotic either alone or more commonly along with betel leaf. Its nut has medicinal and pharmaceutical properties. About 1/10th of world population has a habit of betel chewing. Nut contains polyphenol (11-18 %) which includes flavonols like catechin, epicatechin, leucocyanidin, fat (9-15%), polysaccharides (17-25%), fibre and protein (6- 7.5%) and some minerals like calcium, phosphorous and iron.

Banana is one of the oldest fruits and second largest growing fruit crop in the world. It is also known as the Adam’s Fig and Apple of Paradise. Unlike other fruits banana is an herbaceous fruit plant. Banana breeding started in Trinidad, West Indies in 1922 and in Jamaica in 1924 (Shepherd, 1994). The driving force for this breeding programme was to develop improved Fusarium wilt (Fusarium oxysporum F. sp. Cubense) resistant banana for export trade (Buddenhagen, 1990). In 1960 both the programmes were combined under the Jamaica Banana Board. United Fruit Company also started a small breeding programme in Panama in 1920s. In India hybridization work was started at Central Banana Research Station, Aduthurai, Tamil Nadu in 1949. Important banana growing states are Maharashtra, Karnataka, Kerala, Tamil Nadu, Andhra Pradesh, Orissa, Bihar, West Bengal and Assam. Now a days in some districts of Uttar Pradesh, Harichal cultivar of banana is cultivated at commercial scale. In South India banana is extensively used in all auspicious occasions such as wedding, festival and worship. Banana is a good table fruit, besides, the cultivar Nendran is used for cooking, it is also used for preparation of halwa, sweet and chips etc. Main bud or heart of banana bunches is also used as vegetable in India, Thailand, Philippines, Malaysia and Indonesia. Musa textiles is known for strong fibre quality.

4.1 Introduction

Banana (Musa spp.) is one of the oldest fruits and second largest growing fruit crop in the world. It is also known as Adam’s Fig and Apple of Paradise. Important banana growing states are Maharashtra, Karnataka, Kerala, Tamil Nadu, Andhra Pradesh, Odisha, Bihar, West Bengal and Assam. In South India, banana is extensively used in all auspicious occasions such as wedding, festival and worship. Banana is a good table fruit. The cultivar Nendran is used for cooking; it is also used for preparation of halwa, sweet, chips etc. Main bud or heart of banana bunches is also used as vegetable in India, Thailand, Philippines, Malaysia and Indonesia. Musa textiles is known for strong fibre quality. Ripe banana contains about 20% sugar (but no starch) and about 4.7% protein. Further, it contains vitamins A, B, C, D and E and minerals such as K, P, Ca and Fe.

Ber (chinese date, chinese fig or bore) is an important minor fruit of India. It is hardy fruit tree cultivated all over India and also known as poor man’s fruit. It is most widely cultivated in Punjab, Haryana, Rajasthan, Uttar Pardesh, Madhya Pradesh, Bihar, Gujarat, Maharashtra etc. Ber can do well even under marginal growing conditions and provides quality yields at low cost (Shukla, 1996). Fruits are mostly eaten fresh but the other forms, such as dried, candied, squash etc can also be prepared from ber. Stem bark, root and leaves have some medicinal values. Leaves of ber are used as fodder in dry regions. Ber tree (Ziziphus xylopyrus) is a host plant for rearing the Tachardia laccad a lac insect (Sharma and Kore, 1990).

11.1 Introduction

Ber (also called Indian Jujube, Indian Plum, Chinese Apple and Masau) is an important minor fruit of India. It is most widely cultivated in Punjab, Haryana, Rajasthan, Uttar Pradesh, Madhya Pradesh, Bihar, Gujarat, Maharashtra etc. Fruits are mostly eaten fresh but other forms, such as dried, candied, squash etc. can be prepared from ber. Stem, bark, root and leaves have some medicinal values. Leaves of ber are used as fodder in dry regions.