eChapters

Pomegranate is one of the popular fruit crops of tropical and sub tropical in India and enjoys high reputation for its dietetic and medicinal properties. The fruit is native of Iran and is extensively cultivated in Mediterranean countries like Spain, Morocco, Egypt, Iran, Afghanistan. It is also grown to a lesser extent in Myanmar, China, Japan, India and U.S.A.

The area under pomegranate cultivation has increased several folds in the recent past after the introduction of soft seeded varieties. Out of 30,000 hectares under this crop in India, Maharashtra state alone grows over eighty per cent of area with an average production of 8000 kg ha^-1 (Mote et al.,1992). The export of pomegranate fruits was 1790.3 tonnes in 1991-92 valued at rupees 215.3 lakhs which was more than two fold over the previous year’s export earnings (Chadha, 1994). The utility of the crop is gaining importance and at present new orchards of pomegranate are being planted on large acreage. Because of its versatile adaptability, hard nature, low maintenance cost, steady and high yields, fine table and therapeutic values, better keeping quality and very good export potential, there is greater scope for extending its area and increasing the production especially in the hot and semi arid regions of India.

In pomegranate, market preference is for the attractiveness of the fruit which are exhibited through better size, colour with bold and deep coloured arils besides possessing high juice-content. However consumers prefers the softness of the arils with low acidity. While, there are number of cultivated types available with these desirable characters, there are hardly cultivar having a combination of all or atleast a majority of them. Pomegranate improvement work, therefore has to be oriented towards the production of soft seeded type with other desirable characters (Phadnis, 1974).

The mango is one of the choicest fruit crops of tropical and sub-tropical regions of the world. Its popularity and importance can easily be realized by the fact that it is referred as ‘King of fruits’ in the tropical world. India ranks first, both in area (2.51 million hectares) and production (15.02 million tonnes) of mango and it contributes to 42.1 per cent of world’s mango production. Presently, India is home to more than 1000 mango cultivars differing in taste, aroma, size, fibre content and pulp characteristics.

Though more than 1000 mango cultivars are reported, only 25-30 are of commercial importance. Most of the important cultivars are not amendable to hi-tech cultivation practices and do not meet the requirements of modern horticultural production systems like precocity in bearing, dwarfing, regularity in bearing with high yield, resistant to diseases, pests, physiological disorders and good keeping quality. Hence, breeding attempts to develop an ideal mango variety would be dwarf, regular bearer with medium size fruits (250-300g). Additionally, it should be highly tolerant to various fungal and bacterial diseases. A variety suitable for pickling is also gaining the attention of the breeders. An ideal pickling type should have high acidity, high fibre, good texture and characteristic raw mango flavour.

The first and foremost step in developing an ideotype is to explore the vast diversity and systematic characterization for utilization in the breeding programme.

Considering the importance of conserving the available gene pool, systematic exploration works are being carried out by National Institute like Indian Institutes of Horticultural Research (IIHR, Bangalore), National Bureau of Plant Genetic Resources (NBPGR, New Delhi) and others to collect and conserve the germplasm in Field Gene Banks. Recently, in addition to the traditional germplasm conservation methods, cryopreservation is also gaining importance especially in the conservation of nuclear gene (pollen parent) diversity for future breeding.

Utilizing the conserved germplasm in the breeding programme requires precise information on the genetic relationships within such germplasm. Information on the genetic distance among the germplasm accessions will also help to avoid the duplicates, widen the genetic base of the core collections and ultimately help in preserving the valuable diversity.

Jack bearing the largest edible fruit in the world, is a tropical evergreen tree belonging to the family Moraceae. Previously, it was known as Artocarpus integrifolia.It is an indigenous fruit crop, probably originated in Western Ghats of India (Santapau, 1966) and is widely cultivated in South Asia, East Indies and other warm areas of both the hemispheres. Jack tree is commonly grown in Burma, Malaysia and Brazil. It is hardly recognized as a commercial fruit crop in India even though it is widely distributed in Southern states viz,. Kerala, Tamil Nadu, Karnataka and Andra Pradesh. It is also cultivated in other states like Assam, Bihar, Orissa, Maharashtra and West Bengal. It is an invariable component of Kerala’s homesteads and also grown as a shade crop in coffee, arecanut and cardamom plantations. It is also grown as a standard crop in pepper plantations. The crop has got a rare distinction of being used as the staple food of Kerala tribals and also in some African countries like Uganda.

Gamboge tree Garcinia gummi-gutta (L.), is a member of the Clusiaceae family. It is a botanical cousin of Garcinia indica Choisy. It is a cross pollinated crop. It is called as Kodumpuli in Kerala. It is native of Tropical Asia. It thrives best in the ever green forests of Konkan, stretching from southward to the Kerala coast and Western Ghats upto 1800 m min the Nilgiris (CSIR, 1956). In Kerala, majority of the populations occur naturally in the central districts of Kerala, of different altitudes ranging from sea level to 200 m.

It is one of the potential under-exploited fruit crops. Now, it is gaining momentum for its commercial, industrial and medicinal importance. The dried rind is sold in the market at the rate of Rs. 200 per kg (Muthulakshmi et al.,1999). It is used as a food preservative, spicy source with astringent taste. It is meant for its good flavour, taste and keeping quality. The fruits are too acidic and cannot to eaten raw. The dried rind has unique use in fish curries for imparting its delicate flavour (Sarah, 1998). Dried rind can also be used for polishing gold, silver and coagulating rubber latex. Hydroxy citric acid (HCA) is a very rare natural components being extracted from the rind and is marketed in the name of ‘citrin’ and ‘citrimax’. It has high export value and fetches considerable foreign exchange. In foreign countries, it is used as a herbal medicine against cardiac diseases and for weight management through blocking fat synthesis. The derivatives of acid are potent metabolic regulators of obesity. The unique acid, lowers the blood lipids such as cholesterol and triglycerides by triggering the fatty acid oxidation in the liver through thermogenesis.

Rice (Oryza sativa L.) is the most popular crop and source of food for more than one-third of the world’s population. About 90 % of the world’s rice is grown in Asia and more than 3 billion Asians rely on rice for 30–75 % of their total calories (Braun and Bos 2004). It has been cultivated in four different ecosystems viz., irrigated, rainfed low land, upland and flood-prone (Halwart and Gupta, 2004). Since rice planting has been started around 8,000 to 9,000 years ago, the level of diversity would be more than it expected. The genetic diversity plays a major role in survival and adaptability of a species, as changes happen in environment. Organism needs to make changes in the phenotype that enables it to adapt and survive in unfavorable conditions. The inadequate knowledge on diversity of a species reduces the chance of widening the genetic base of that species. The venture of green revolution in Indian agriculture has narrowed down the genetic base of any popular crops. Therefore, collection, conservation and utilization of germplasm are very much needed in the present scenario for improving the varieties against biotic and abiotic stress.

Morphological and biochemical trait based diversity studies being largely influenced by environment as phenotype and/or enzymes are altered by the environmental and developmental changes. Therefore, adapting DNA based molecular markers for genetic diversity studies would be more reliable to overcome the environmental influence. Being molecular markers as a versatile tool, it establishes its strong holds in several fields of life science. Molecular marker based technique does not require DNA sequencing but it is generally based on nucleic acid hybridization or polymerase chain reaction (PCR). The PCR based tests has several advantages over hybridization technique as it is cheaper, speedy and sensitivity. Therefore, PCR based markers have been used extensively in genetic diversity study for assessing genetic variation within the species. The recent advent of molecular and computational tools now enables the estimation of genetic diversity and allele frequency between the genotypes easy.

A successfully developed new cultivar should have stable performance and broad adaptation over a range of environments, seasons and locations, in addition to high yield potential. Evaluating performance stability and range of adaptations is becoming increasingly important in breeding programs. Since stability and adaptability are important selection criteria in breeding programs, in depth research on stability is needed for a better estimate of crop performance and adaptability (Lin and Binns, 1988).

Having realized the importance of genotype × environment, several biometrical models have been developed to analyse the stability of genotypes. Such models have been discussed, critically reviewed comprehensively by Knight (1970), Freeman (1973), Hill (1975), Brain (1986) and Gautam et al. (1986).

However, the various methods proposed for the statistical analysis of genotype × environment are based on one of following approaches (Jain, 1982). In the first the variance interactions are estimated by equating the observed mean squares in the analysis of variance to their expectations on the random model. The second approach called phenotypic approach by Hill (1975) used the technique of partitioning the genotype _× environment component of variability into its linear and non-linear portions of assessing the stability of genotypes over a range of environments. This approach which is very widely adopted in plant breeding, came to be known as “Joint regression analysis”.

Many workers contributed for the development of this approach, notably Finlay and Wilkinson (1963), Eberhart and Russell (1966), Freeman and Perkins (1971) and Shukla (1972). The third approach called genotypic approach which also suggested by Hill (1975) is based on fitting of general models in which it specify, the contribution of genetic, environmental and genotype × environment to the generation means and variance. The contributions for he development of this approach were from Bucio (1966), Bucio and Hill (1966), Bucio et al. (1969), Perkins and Jinks (1968), Breese (1969), Hill and Perkins (1969) and Perkins (1970). Linear regression and joint regression models were more commonly used due to their simplicity and reliability in biological experiments. In general, these approaches fall conveniently into two parts.

Stability and adaptability studies have been carried out for upland or hill rice varieties also. Amirthadevarathinam (1987) observed significant genotype × environment and reported that both linear and non-linear components were equally important for assessing stability of yield and four yield components in four environments. Both TKM 6 and Poongar were high yielding with adaptability and stability in addition to being especially suitable for unfavourable environments. Kuruvaikalangium was the best performer for all characters except grains per panicle for which PM 1023 was the best.

Dineshkumar (1987) evaluated sixteen early maturing genotypes for their stability in four major agroclimatic zones of the Karnataka state. No genotype was found stable as they observed more of unpredictable variance (pooled deviation) than predictable (genotype × environment linear).

Ganesh and Soundarapandian (1987) observed that stability of number of ear bearing tillers was a major contributor for yield stability Shivaprakash (1987) reported similar results.

Studies by Sinha and Biswas (1987) on yield and yield components of twenty photo insensitive rice cultivars revealed IR-36 and IET-4094 as high yielder with unit ‘bi’ and least ‘S² d_i’ and hence they were recommended to be suitable for general cultivation for all the environments under study. Similarly IET-2881, which had high mean yield with high ‘bi’ was found suitable for high yielding environments.

Genotypes × Season interaction is of major importance to the plant breeder in developing stable high yielding varieties of crop plants. When new varieties are tested over a series of seasons, the relative ranking of the varieties for any given attribute is rarely the same at each season. This increases the difficulty of identifying superior stable genotypes. Comstock and Moll (1963) have shown statistically the effect of large genotype × environment interaction is reducing the progress from selection.

In order to minimize the genotype × season interaction, stratification of the seasons can be practiced. The implification in stratification is that suitable selections could be made for the infavourable and favourable seasons encountered within the sampling units proposed. Stratification of environments has been practicised for a long time to identify suitable selection that could be made for the narrower range of environments encountered within the sampling units proposed. However, even with this refinement of technique, the interactions of genotypes with locations in a sub-region and within environments encountered at the same location in different years, frequently remain too large (Allard and Bradshaw, 1964). Little is known about the environmental factors contributing to such interactions. Sprague (1966) suggested that even if such information are available, the possibility of materially reducing genotype × environment interaction in field experiments would remain questionable.

On line with Allard and Bradshaw’s (1964) statement, the interactions of genotypes with locations in a sub-region with different seasons at the same location in different years, may remain too large. As far as sesame is concerned, practically nothing is known about the factors contributing to such genotype × season interactions. Hence, the progress in breeding a stable variety of sesame has been slow. Since sesame in India is grown essentially as rainfed crop, fluctuations in yield levels closely follow changes in the environment and seasons. Breeding for multiseasonal variety for may small regions within a state is being considered necessary. There is widespread awareness of achieving stability of production as well as improvement in yield of sesame under low and high-in-put conditions as well as for different seasons.

In developing countries, the most important challenges is to increase food productivity for the continuously growing population from limited land area. To great extent, this challenges has been met with good amount of success in cereals but not in case of pulses. Pulses from an indispensable component of Indian agricultural too. Further, it is an excellent source of protein to the growing population who are mostly vegetarians. The high level of lysine in the protein makes them ideal supplement to cereals. Moreover, they are endowed with nitrogen fixation, and thus improve the soil fertility.

India is one of the major pulse growing countries in the world with a total area of 23.31 million hectares, with a total production of 14.50 million tones (Source: FAO, 2011). The area, under pulses has decreased particularly during the last decades as a consequence of intensive diffusion of techniques in green revolution on the high yielding varieties of cereals. This is mainly due to the low yield potential of legumes even under high input conditions.

Among the various pulses, blackgram/urd bean known scientifically as Vigna mungo (L.) Hepper, former known as Phaseolus mungo (L.) belonging to the tribe Paseolea of family Papilionaceae is of immense importance. It is the most widely grown and highly esteemed grain legume. In India it occupies 2.9 million hectares with an annual production of 1.24 million tones (Source: FAO, 2011).

In Asia this crop is widely cultivated in India, Pakistan, and Bangladesh both in kharif and rabi seasons. In India, it is mainly grown in the states of Madhya Pradesh, Maharashtra, Uttar Pradesh, Rajasthan, Karnataka and Bihar. Tamilnadu is placed eleventh and tenth in area and production respectively (FAO, 2011).

Ramya (2012) studied stability of 12 hybrids in three environments. The researcher selected the 12 hybrids based on the standard heterosis. The hybrids are: P₂ × P₅, P₁ × P₅, P₃ × P₅, P₂ × P₇, P₂ × P₃ and P₁ × P₂. Among the twelve hybrids, the cross combinations viz., P₂ × P₅, P₂ × P₇ and P₂ × P₃ showed non-significant reciprocal effects for most of the traits including fruit yield per plant. The direct hybrid P₁ × P₂ showed less than 40 per cent standard heterosis for fruit yield per plant. It also exhibited low standard heterosis for number of fruits per plant. Eight hybrids viz., P₂ × P₅, P₅ × P₁, P₃ × P₅, P₂ × P₇, P₁ × P₅, P₂ × P₃, P₂ × P₁, P₅ × P₃, based on standard heterosis (>40 per cent) and their parents viz., P₁, P₂, P₃, P₅, P₇ were selected and grown in three different locations for finding their stable performance. Several authors have suggested different types of models to identify the stable genotypes. The model most commonly preferred by breeders is Eberhart and Russell (1966) model which was used in the present investigation. Various stability parameters were worked out on the available data and the results were discussed below.

Among the ten traits studied, days to first flowering, fruit weight and fruit girth showed non-significant genotype × environment interaction. Hence, the remaining seven traits were taken for further analysis. ANOVA across environment revealed highly significant mean sum of square for genotypes, indicating the presence of wide genotypic variability among the genotypes involved in the study for all the characters studied. High significant differences were observed for environments for the traits viz., number of branches per plant, fruit length and fruit yield per plant, indicating the divergence among the growing environments and differential response of genotypes to the various environments. The main factors contributing to the divergence of the environments might be temperature differences during crop season, intensity and duration of sun shine hours. Therefore, the genotypes must be tested over an extensive range of environments for proper assessment of stability of genotypes (Salesh Kumar Jindal et al.,2008).

WORLD HISTORY

Work on ornamental crops was first initiated in Krelage Nursery founded by Ernst Heinrich Krelage for bulbs, lilium and dahlias in 1811. During 1895, Taiwan Agricultural Research Institute was established for ornamental crops. The first yellow corn Golden Bnatam was listed in Burpee’s Catalogue in 1902. In 1909 Primadonna F₁ hybrid variety of begonia was released by the Benary Seed Company in Germany and in the same year Atlee established seed farms at Floradale in California, Erfurt in Germany Anglia in UK and Kashmir valley in India for annual garden flowers. Luther Burbank did excellent breeding works on ornamental trees, flowers and fruits in 1926.

David developed the first successful hybrid in Nasturtium (1934) and Red and Gold hybrids of marigold (1937). In 1940, Burpee was successful in chromosome doubling and transformation of wild flowers and developed “Bright Scarlet” and “Super Tetra Series” in antirrhinum and “Jumbo Scarlet” in zinnia. He produced new strains of marigold, zinnia, nasturtium and snapdragon. William Houston in 1940 initiated the Ageratum breeding. Morden Research station was set up in Western Canada in 1945 for ornamental crop breeding. In 1968, Dr. Gustav A.L., Mehlquist was recognized as an international authority on breeding of ornamental plants. George J. Ball breeder of peas, asters and calendulas was a pioneer for the supply of dwarf impatiens and orange impatiens ‘Super Elfin’ in 1968. Israeli breeders during 1990 gave more attention towards the development of carnation, lily and anemone varieties. Longwood Gardens in 2008 got success in breeding of stock. The first record of chrysanthemum hybridization, germination and selection of superior seedlings occurred in 1827 by M. Bernet.

INTRODUCTION

Various procedures which are used for genetic improvement of crop plants are known as breeding methods. The breeding approaches that are more commonly used for genetic improvement of crop plants are called conventional breeding methods. Following conventional breeding methods were followed in improvement of crops.

(A)Plant introduction

(B)Pureline selection

(C)Mass selection

(D)Progeny selection

(E)Synthetic and composite breeding

(F)Pedigree method

(G)Bulk method

(H)Backcross method

(I)Heterosis breeding

(J)Clonal selection

MUTATION BREEDING

Mutation refers to sudden heritable change in phenotype of an organism or permanent change in the number, kind and sequence of nucleotides in genetic material, while individuals showing these changes are known as mutants. Mutation was first discovered by Wright in 1791 in male lamb which had short legs. Hugo de vries coined the term mutation. In crop improvement the breeders are always look for wide range of variability and stabilization of desirable traits. Mutation breeding is very useful in inducing new variability, which is an essential requirement of any plant breeding programme. The modern day cultivars are the resultants of various breeding techniques like selection, hybridization, tissue culture and mutagenesis including some biotechnological tools. Novelty in commercial traits like colour, size, shape, length, post harvest life of flowers etc. are always valued and preferred by consumers. Mutation breeding by exposing planting material with irradiation has play vital role in inducing novelty in a number of flower crops.

TYPES OF MUTATION

Based on occurrence

(1)Spontaneous mutation: Occur spontaneously in nature and not present in sufficiently high frequency and spectrum.

(2)Induced mutation: Artificially produced by the treatment of various mutagenic agents and present in sufficiently high frequency and spectrum

MOLECULAR BREEDING

Molecular breeding is the application of molecular biology tools, often in plant breeding.

Genetic markers

Genetic marker is defined as a gene whose phenotypic expression is usually easily discerned, used to identify an individual or a cell that carries it, or as a probe to mark a nucleus, chromosomes or locus (King and Stansfield, 1990). Three broad classes of genetic markers are morphological markers, biochemical markers and DNA or molecular markers.

(1) Morphological markers

Morphological markers are based on the visually assessable traits viz., morphological and agronomic traits. However, they are limited in number.

Features of morphological markers

i.They are easy to identify, there is no need of any complex analysis. Trait of interest can be identified visually.

ii.Morphological markers are inherited qualitatively or quantitatively.

iii.Frequency of availability of morphological markers is less in number i.e., morphological markers may not be available for all the traits of interest.

iv.They are influenced by environment.

v.Since they are influenced by environment, they are unstable in inheritance.

vi.They are stage specific i.e., a particular marker will be useful to identify in a particular stage of plant and not any stage of the plant.

vii.They are influenced by genotype and developmental stages of the plant.

Besides drawbacks, the morphological markers are useful to the plant breeders for the selection purpose in certain situation.

INTRODUCTION

Hybrid crops are derived from an established and well-proven breeding method used in ornamental sectors. The development of hybrid crops has been one of the major factors behind a dramatic increase in global crop yields. Hybrid is produced by crossing between two genetically dissimilar parents. Pollen from male parent called pollen parent will pollinate, fertilize and set seeds in female called seed parent to produce F₁ hybrid seeds. For production of a hybrid, crossing between two parents is important, the crossing process will results in heterosis or hybrid vigour. Heterosis is defined as the increase in size or vigour of a hybrid over its parent. This boost in performance, combining the best yield, quality and agronomic characters from each parent, is the result of hybrid breeding.

F₁ hybrid seed offers to the growers a significant benefits in terms of yield improvement, agronomic performance and consistency of end-use quality. But attempting to farm-save hybrid varieties will result in segregating populations producing highly variable offspring, which can lead to reduced yield, disease resistance and quality. As this factsheet explains, it doesn’t make economic or agronomic sense to save seed of an F₁ hybrid. In self pollinated crops it is difficult to cross but in cross pollinated crops it is easier. In nature to create genetic variability and for its wider adaptation in different environmental conditions, flowering plants has adopted many mechanisms for cross pollination. Cross pollination results in genetic heterogeneity and show wider adaptations. F₁ hybrids are in immense usage throughout the world with the rapid increasing interest in flowers, however the total number of the hybrids developed in flowers is still less. In India hybrid seed production is taken up in Bengaluru, Pune, Delhi, Punjab, Haryana, Jammu and Kashmir and Himachal Pradesh for export.

F₁ hybrids in ornamental crops were known even before Shull 1911 when he propounded the classical theory of hybrid vigour in plant breeding for the first time. In 1909 F₁ hybrid variety Prima Donna in begonia was released by Benary Seed Company in Germany. The first commercial F₁ hybrid in petunia was produced by Japan during World War II, in 1942. From 1950 onwards, F₁ hybrids were available in ornamental flowers from The Netherlands, Denmark, Germany, USA and UK besides Japan and produced in flower crops like ageratum, anemone, gerbera, primula, petunia, cyclamen, marigold, pansies, begonia, geranium, portulaca, dianthus, balsam, stock, wall flower, ornamental sunflower, gazania, hollyhock, calceolaria and zinnia. Flower breeders became familiar with the F₁ hybrid production of those crops also which were formerly propagated by vegetative parts such as pelargonium and gerbera. A few important hybrids were first released in petunia in 1940-50, geranium single in 1960, antirrhinum, marigold, pansy and zinnia in 1965, ageratum in 1966, geranium double in 1970, dianthus, impatiens, portulaca in 1976-77, gerbera in 1980 and carnation in 1981.

Intellectual Property Rights (IPR) are legal rights, which result from intellectual activity in industrial, scientific, literary and artistic fields. These rights safeguard creators and other producers of intellectual goods and services by granting them certain time-limited rights to control their use. Protected IP rights like other property can be a matter of trade, which can be owned, sold or bought. These are intangible and non exhausted consumption. The term “intellectual property” denotes the specific legal rights which authors, inventors and other IP holders may hold and exercise and not the intellectual work itself (Chadha, 2013). Intellectual property rights are essentially territorial in nature so, where registration is necessary, for patents, registered designs and trademarks, these will not exist in other countries unless registration has been sought and obtained there.

IP protection is crucial for a sustainable contribution of plant breeding and seed supply. Breeding new varieties of plants, which are developed after contributing number of years to the selective inheritance of traits which provide improved yields, higher quality and better resistance to such plant varieties requires a substantial investment in terms of skill, labour, material and economic resources. However, a new variety once released could in many cases be readily reproduced by others so as to deprive its breeder of the opportunity to benefit adequately from the investment made. Thus, an effective system of plant variety protection (PVP) is a key enabler for investment in breeding and the development of new varieties of plants or improving existing plant varieties, encourage importation of foreign varieties, promote exportation of plant varieties provide access to information of the created products and the methodology of creation for the enhancement of social welfare and generally benefit the market place. An international system of IPR protection for plant varieties expands these benefits by facilitating access to new varieties created in other states (Gautam et al., 2012).

Intellectual property is usually divided into two branches: (i) industrial property and(ii) copyright and the rights which neighbour upon copyright. The principal categories of industrial property are: patents, trademarks, geographical indications, industrial designs and trade secrets. Industrial property according to the 1883 Paris Convention for the Protection of Industrial Property is to be understood in the broadest sense and to apply not only to industry and commerce proper but also to agricultural and extractive industries and to all manufactured or natural products including wines, grain, tobacco, leaf, fruit, cattle, minerals, mineral waters, beer, flowers and flour.

Alstroemeria belongs to the family Alstromeriaceae. Alstroemeria flowers third in terms of popularity and foreign exchange earnings in Kenya after roses and statice. Alstroemeria is named after a Swedish botanist “Baron Klas von Alstroemer”. The popularity of this flower is still growing in the global perspective. It is one of the quickest growing flowers with varying colours like purple, lavender, salmon, red, pink, white, orange, yellow and bicolour. It is grown for cut flowers but is also popular as a potted plant. It is grown in the higher ranges of the Western Ghats, Himachal Pradesh, Jammu & Kashmir and Karnataka in India.

ORIGIN AND DISTRIBUTION

The alstroemeria, sometimes called the Peruvian Lily, is native to Tropical America particularly Chile. There are some 30 species of alstroemeria that occurs in grasslands and Pampas of South America, Argentina, Bolivia, Brazil, Chile, Paraguay, Peru and Venezuela. In America, it flourishes in different climates from high in the Andes mountains to the deserts of Pacific coastline. Even deep into the tropics of Brazil, the plants flourish abundantly. It was introduced to Europe in 1753 when seeds were sent to Europe by Claus Alstroemeria of Sweden during his travels in South America. It was widely cultivated in England during the early 1800’s and it was John Hedgewood, the founder of the Horticultural Society who first introduced the white variety in 1832. Since then, the Alstroemeria has been hybridized extensively. It is named after a Swedish botanist, Baron Klas von Alstroemer, who brought the first seeds of A. oelegrina from South America to Europe. Columbia is the number one in the world as far as growing area of alstroemeria is concerned. The second and third positions are taken by the Netherlands and Japan respectively. Other producing countries are United Kingdom and Italy. Alstroemeria is a new introduction into the world floriculture scene.

Most commercial cultivars of alstroemeria are patented and protected through plant breeder’s rights. Because of the fact that quite a large number of new varieties and hybrids of alstroemeria are planted in various countries in each year, royalty is paid based upon their square footage under production or number of plants leased. The new Inca Collection bred by Konst Alstroemeria B.V. is a breakthrough in the presently dwarf alstroemeria assortment. The new extended range of alstroemeria cultivar offers the opportunity to grow these top quality perennials in the garden, pots and for cut flower. These dwarf type plants reaching a height from 20-50 cm, combine with the quality and colours of the long lasting flowers. New varieties have gained popularity in California (USA), New South Wales (Australia) and Spanish south east coastline, Sicily (Italy). However, for a very long time already alstroemeria has been cherished worldwide for its exotic flowers.

In India cultivation of alstroemeria has been started around Bengaluru, Pune and Hyderabad. It has been introduced in Palampur, Solan, Srinagar and hills of Uttrakhand and Ooty, to popularize this crop among growers for domestic and export markets (Singh, 2006).

Anthurium (Anthurium andraeanum Linden) belongs to the complex family Araceae, is a slow growing perennial that requires shady, humid conditions as found in tropical forests. The name anthurium is derived from Greek word “anthos” meaning flower and “oura” tail referring to the spadix. Thus, anthurium is also known as “tail flower” (Tajuddin and Prakash 1996). Anthuriums are gaining popularity due to higher returns per unit area and their beautiful and attractive long lasting flowers. Anthurium ranks ninth in the global flower trade and commands a respectable price both for its cut flower and whole plant. Commercial production has focused on two major species viz. A. andraeanum and A. scherzerianum. A. andraeanum is grown mostly for cut flower production, the main production areas being Hawaii, The Netherlands, and some other tropical and subtropical countries. A. scherzerianum is sold as a flowering potted plant, with main production areas located in Europe. In India, the anthurium cut flower industry is still in its infancy. At present anthuriums are mostly grown in some small gardens and nurseries. The important states cultivating anthuriums are Assam, Kerala, Tamilnadu (Salem), Karnataka (Coorg) and Mizoram where the favourable climate exists (Agasimani et al., 2011).

ORIGIN AND DISTRIBUTION

Anthurium andreanum Linden Ex Andre is named after a French Horticulturist Edouard Francois Andre (1840-1911) who discovered it in 1876 and described it for the first time. The genus includes around 1000 species (Croat, 1980). Most of them are tropical herbs originating from warm areas of Central and South America (Gantait et al., 2008). The distribution of this genus extends from northern Mexico and the Greater Antilles to southern Brazil, northern Argentina, and Paraguay (Croat and Sheffer, 1983). Most anthurium species are native of tropical rain forests and are primarily epiphytic in nature. Thus, in their natural habitat, they receive ample, frequent water with good drainage. In Brazil, it is noted the occurrence of around 130 native species (Gonçalves, 2003).

In Malaysia, anthurium is mostly grown in Pahang (Cameron highlands), Selangor, Perak and Johor mainly for cut flower production. The Netherlands is the chief supplier of the anthurium in the European market. Hawaii covers 60% export share of Japan and the rest of the west coast of the USA. The Caribbean produces the anthurium for the east coast of United States and Canada. Mauritius is one of the important producer of anthurium and it exports to Japan and Italy. While some eastern countries like Taiwan, Philippines, China and Thailand mainly produce for their local market. In India, anthurium is commercially grown in Kerala, Karnataka, Tamil Nadu, West Bengal, Maharashtra and north east India. Kerala is identified as the most desirable place for growing anthurium (Singh, 2006).

Antirrhinum is a genus commonly known as snapdragon belongs to family Scrophulariaceae. Snapdragon is one of the flowering plant present with vast genetic diversity in India. The term Antirrhinum is derived from the Greek word “anti” means like and “rhin” means a nose, referring to the snout like shape of the flower. The cultivated Antirrhinum majus is grown in tropical, subtropical and temperate countries. The common name snapdragon, originates from the flowers reaction to having their throats squeezed, which causes the mouth of the flower to snap open like a dragon’s mouth. Flowers are borne on terminal long spikes of many colours except blue and with numerous shades. It is one of the excellent cut flowers which have long lasting qualities. These are also used as garden plants, bedding plants, in rockeries or herbaceous borders and as potted plants. Plants can be grown for cut flowers in an open field or under protection depending on the climate. The leaves and flowers are antiphlogistic, bitter, resolvent and stimulant (Chiej, 1984).

ORIGIN AND DISTRIBUTION

The garden snapdragon (Antirrhinum majus) has emerged as a model organism during early studies of inheritance and mutation (Darwin, 1868) because of its diploid inheritance, ease of cultivation and variation in morphology and flower colour. The genus Antirrhinum includes 15 New World species in the section Saerorhinum and 21 Old World species in the sections Antirrhinum and Orontium. A. majus (Snapdragon) is a species of Antirrhinum native to the Mediterranean region, from Morocco and Portugal north to southern France and East to Turkey and Syria. Most species are found around the Mediterranean Sea and in North America (Stubbe, 1966). The progenitor of the modern snapdragon is from the Mediterranean region and more specifically, it is native to southern France. It was believed that the snapdragon reached Britain with the Romans at an early date, where it has been naturalized on mountainous regions. Thereafter, it spread to different parts of the world from Britain.

The scientific name of the genus Antirrhinum was first defined by Carl von Linne in the year 1753. In the year 1914 Professor Erwin Baur the founder of the MaxPalnck-Institut für Züchtungsforschung in Cologne published a volume called ‘Einführung in die experimentelle Vererbungslehre’ (Introduction to Experimental Genetics) 2nd edition. This book contained several plates with reproductions of water colour paintings of snapdragons as examples. He crossed a white line with a red line resulting in a pink flower. This pink flower then was selfed and the offspring showed the typical distribution of 1 white, 2 pink and 1 red as a demonstration of Mendelian inheritance. Antirrhinum glandulous is native of California. In United States of America, winter flowering cultivars were introduced in 1926 by Frank Volz of Cheviot, Ohio. In 1959, snapdragon were included in the Agricultural Censuses of Horticulturally Special Crops and ranked seventh as a cut flower crop.

The genus Antirrhinum offers an inexhaustible resource for genetics-based studies in developmental biology, biochemistry and evolution (Stubbe, 1966). Wide diversity of Antirrhinum species, combined with classical and molecular genetics, the two traditional strengths of Antirrhinum provides an opportunity for developmental, evolutionary and ecological approaches (Schwarz-Sommer et al., 2003). Transposons responsible for flower variegation were identified in the 1980s. The Max-Planck-Institute in Cologne, Germany, allowing genes involved in flower and leaf development and in pigmentation to be isolated by transposon tagging. Population genetic studies that have been applied to Antirrhinum species show different population sizes, geographic distributions and breeding systems, from self-fertility to obligate out-crossing (Jimenez et al., 2002 and Mateu-Andres and de Paco, 2006). It was introduced to India with the advent of British. Among them, only A. majus has been domesticated as an ornamental.

Carnation (Dianthus caryophyllus L.) is a member of the family Caryophyllaceae. Dianthus word was first coined by Theophrastus from Greek word “dios” meaning divine and “anthos” meaning flower. It is a half hardy, perennial plant, propagated commercially by terminal cuttings. It is being produced as a cut flower in the world. Carnation cut flower trade contributed about 15% of the world flower market. Till now, new carnation varieties have been produced mainly through conventional breeding method. The breeding procedure typically consists of hybridization, self-pollination and selection (Holly and Baker, 1992). The breeding programme did not get enough momentum due to heterozygosity, limited gene pool and inadequate knowledge of genetic make up. However genetic engineering and molecular breeding have opened the way to achieve novel colour and good vase life of flowers.

ORIGIN AND DISTRIBUTION

It is native of Mediterranean region and its distribution is restricted to Greece, Italy, Sicily and Sardinia (Tutin et al., 1993). According to Pliny, the centre of origin of carnation is Spain. The generic name Dianthus comes from writing of the Theophrastus who lived about 300 BC; dios, divine and anthos, flower; the flower of the gods. The species name caryophyllus is coined from Greek word Caryon, nut and phyllon, leaf. Linnaeus chose the species name, caryophyllus, after the genus of clove, as the fragrance from carnation is reminiscent of clove. The common word carnation probably comes from a Greek word coronation because of its use as decorating the crowns of Greek athletes during ancient times. Carnation has been cultivated for over 2000 years and today commercial cultivation is the result of 200 years of improvement and breeding. During sixteenth century its improvement work was started in various countries. The gardens of Italy, France, Germany, Holland and England have contributed much to the development of cultivars. The natural climates for carnation occur near 30^o N or S latitudes of equator and on the western edges of the continents.

Gerbera (Gerbera jamesonii), a tender, stemless, perennial flowering plant, belongs to family Asteraceae. Gerbera was named in honour of the German botanist and naturalist “Traugott Gerber” who was a friend of Carolus Linnaeus. Gerbera is very popular and widely used as cut flower or as a decorative garden plant. It was discovered in 1878 near Barberton, eastern Transvaal (South Africa), therefore, referred as the Barberton daisy or Transvaal daisy. It is ideal for beds, borders and pots and is the fifth most important cut flower in the world.

ORIGIN AND DISTRIBUTION

Gerbera is originated from South Africa. The native distribution of genus Gerbera comprising 30 species, extends to Africa, Madagascar, tropical Asia and South America (Hansen, 1985 and Bremer, 1994). Whereas, it was also reported that gerbera is found in the temperate and tropical parts of Asia and Africa and about 7 species are recorded in India. This plant is distributed in the temperate Himalayas from Kashmir to Nepal. Gerbera is commonly seen on the dry slopes near Shimla and on hillsides above Gulmarg in Kashmir (Anon., 1956). The G. jamesonii is found naturally in grassland in sandy soils in Mpumalanga and Barberton district. The genus Gerbera has approximately 14 southern African species, some of which are found only in Western Cape. G. aurantica is endemic to the mist belt grassland region of KwaZulu-Natal and is known from approximately 9 scattered populations from Helahela in the south to Babanango in the north. This species is under threat of extinction due to habitat fragmentation and degradation. In India they are distributed in temperate Himalayas at altitudes of 1300-3200 metres and observed from Kashmir to Nepal.

The genus Gerbera is named in honour of German naturalist Traugott Gerber who travelled in Russia in 1743 and started first botanical garden in Moscow. However, for species i.e. jamesonii, it is a matter of different opinion. The jamesonii was named for Dr. L. S. Jameson, a British colonial statesman in South Africa who was one of the discoverers of this species, whereas in some literature, the species was named after Robert Jameson who collected live specimens while on a prospecting expedition to the Barberton district in 1884, even though the species collected on three earlier occasions by other people. In 1888, Medley Wood, the curator of the Durban Botanical Garden sent plants to Kew, which subsequently flowered. This is native to South Africa, Madagascar, Asia, South America and Indonesia.

The first official description of South African species G. jamesonii was made by J.D. Hooker in Curtis Botanical magazine with coloured illustration in 1889. An active society on this subject is to be found in Australia under the name W.A. Gerbera Society. The society has used a classification system of gerbera. The breeding of gerbera started at the end of the nineteenth century in Cambridge, England, when two South African sp. were crossed. G. jamesonii native to Natal and Transvaal and G. viridifolia from western Cape were crossed together by R. Irwin Lynch in 1887, after both these species were introduced to Cambridge Botanic Gardens by Jameson and the hybrid was named as G. x cantabrigensis.

Gladiolus (Gladiolus spp.) belongs to family Iridaceae and subfamily Ixioideae. Gladiolus is a Latin word, the diminutive of “gladius” meaning sword, sometimes called as sword lily. The plant is herbaceous perennial and semi-hardy. It is commonly grown from rounded, symmetrical corm, which is enveloped in several layers of brownish, fibrous tunics. The cultivated gladiolus is a complex assemblage of cultivars originating from complex ancestry of natural selection or hybridization involving several species. Modern garden gladiolus is a complex of at least twelve species (Raghava, 2000). It is an important commercial cut flower in both domestic and international markets.

ORIGIN AND DISTRIBUTION

There are two main centres of distribution of Gladiolus species, one in Mediterranean basin and the other in southern and central Africa (Lewis et al., 1972 and Delpierre and Du Plessis 1974). Most of the species are native to Mediterranean region and tropical parts of the South Africa, particularly the region of the Cape of Good Hope (Raghava, 2000).

The name gladiolus was coined by Pliny the Elder (A.D. 23-79) to describe the shape of the leaf which resembles that of sword (Latin word gladius meaning sword). It is native to South Africa. Most of the species scattered throughout South Africa, Zambesi River, Rhodesia, Mt. Kilimangero, Tanganyika, Natal and Cape of Good Hope, Republic of South Africa, Madagascar and Abyssinia (Ethiopia). Gladioli are said to be cultivated since the days of ancient Greece. As evidenced by a record in Lyte’s Nieeve Herball, it is known since 1578, first introduced in France and soon after spread to England, Germany, Holland and North America. No evidence is available that any species of gladiolus of South African origin was ever grown in Europe prior to 1700. Cornutus described Gladiolus aethiopion bearing light red-scarlet flowers and is from Cape region. In the middle of eighteenth century the South African species were introduced into Europe. Gladiolus tristis closely related to G. grandis was first introduced in Chelsea Physii Garden, UK in 1745 from Cape region and Walat. G. illyricus is native to Mediterranean region. In 1749, Gladiolus grandis was naturalized in Spain from South Africa.

By 1816, France, Holland and Belgium started raising hybrids, the first being Ghant Gladiolus, Gladiolus gandavensis, in 1841, by crossing G. cardinalis and G. psittacinus and vice-versa. Gladiolus primulinus having central petals with hooded characteristics was collected from its native place on the banks of Zambesi River near Victoria Falls in 1902. Victor Lemoine of France used G. primulinus for producing ‘Primulinus Hybrids’ during 1906-1908 while A.E. Kunderd of Indiana, U.S.A. in 1915 produced a similar race ‘Primulinus Grandiflora Hybrids’ by using same parentage as Lemoine. In 1907, first ruffled gladiolus with fimbriated and frilled petals was produced by the efforts of Kunderd.

Gladiolus is a potential flower crop for floriculture industry in India. Gladiolus is commercially grown in West Bengal, Maharashtra, Uttar Pradesh, Uttarakhand, Punjab, Haryana, Sikkim, Jammu and Kashmir, Karnataka, Gujarat, Himachal Pradesh, Tamil Nadu, Madhya Pradesh, Delhi and Rajasthan. The area of the states of U.P., Uttarakhand, Haryana, Punjab, Rajasthan and H.P. and surrounding Delhi grow gladiolus mostly for Delhi flower market (Singh, 2006).

Heliconia belongs to family Heliconiaceae is popular as ornamental plants and cut flowers because of their brilliant colours and exotic appearance. The genus name “Heliconia” is derived from “Helicon”, a mountain in southern Greece regarded by the ancient Greeks as the home of the Muses, thus suggesting the relationship between these plants and the bananas, genus Musa (Berry and Kress, 1991). Unique features of the Heliconiaceae are (1) medium to large erect herbs rising from underground rhizomes;(2) each erect shoot is composed of a stem and leaves, whereas the stem is made up of an axis covered by overlapping sheathing leaf bases, technically called a pseudostem;(3) inverted flowers and (4) the presence of a single staminode. Heliconia has long been popular horticulturally because of their showy inflorescences. It was so attractive that early explorers of the tropics returned to Europe with several species that became prized green house specimens. Heliconia, grown in nearly all of the tropical regions of the world, including Africa and Asia. In India, heliconia is grown in North-East region and Kerala.

ORIGIN AND DISTRIBUTION

Heliconias are native only to Central and South America and some of the islands of the South Pacific, for horticultural and commercial popularity. Berry and Kress (1991) stated that they are native from Tropical America and found at different altitudes, from sea level up to 2,000 meters in Central and South America, and up to 500 meters in the South Pacific Islands (Criley and Broschat, 1992).

Most of the Heliconia species are distributed primarily in Neotropical areas from the North of Mexico to the South of Brazil (Santos, 1978, Dahlgren et al., 1985 and Kress, 1990). A small Paleotropical group, about eight species, occurs in islands of the South Pacific (Tomlinson, 1969 and Kress, 1985). There are approximately 40 Brazilian species with two primary areas of distribution: the Amazon basin and the Atlantic coastal forest (Kress, 1990). About 100 or more species, native mainly to tropical and subtropical South and Central America (Cronquist, 1981).

Several attempts have been made over the years to sort out the taxonomy of the Old World Heliconia. Baker (1893), followed by Schumann (1900) and Winkler (1930), dealt with the problem of identity and origin of these taxa by assigning them to the neotropical species Heliconia bihai L. Others such as Ridley (1908a, b), Backer (1920) and Green (1969) recognized the distinctive and endemic features of the paleotropical taxa that separated them from the neotropical species. This ornamental rhizomatous herbaceous plant from the Heliconia genus, belongs to the Musaceae family, now constitutes the Heliconiaceae family in the Zingiberales order. The various species of Heliconia are subdivided into five subgenera: Heliconia, Taeniostrobus, Stenochlamys, Heliconiopsis and Griggisia; and 28 sections (Kress et al., 1993).

Kangaroo Paws (Anigozanthos spp.) belongs to family Haemodoraceae. It is commonly known as Sword-like or Iris-like. The word Anigozanthos was coined by French botanist Jacques de La Billardiere. This is derived from Greek words “anoigo” meaning to “expand or unequal” and “anthos” meaning “flower or irregular corolla”. It can be used as cut flower, pot plant, landscape plants or dry flower (Gui et al., 2005). The kangaroo paws is gaining popularity among gardeners, horticulturists, nursery man also in world market due to its beautiful, unique, colourful pattern of flowers and also being perennial in nature.

ORIGIN AND DISTRIBUTION

It is originated in Australia. Louden’s Hortus Britanicus, A Catalog of a Plants Endemic, Cultivated or Introduced to Britain, written in 1830, states that in 1808 Anigozanthos manglesii was introduced by Robert Mangles who had possibly received seed from his brothers in New Holland. Anigozanthos rufus was introduced into cultivation in 1824. The Royal Horticultural Society Dictionary of Gardening dates of the Botanical Magazine Plates are listed as A. flavida 1808 (BM 1151), A. manglesii 1833 BR, A. pulcherrimus 1844 (BM 4507). The first report of introduction of 8-10 species of Anigozanthos under green house from Australia in USA was reported in 1902 edition of Bailey’s Cyclopedia of American Horticulture. The first hybrid Bush Gems was introduced from Australia by Mr. Meyer, he had also made good selection of A. rufus and A. pulcherrimus these were propagated by Oglesby tissue culture lab. Many of them were released for sale at the Huntington Gardens annual plant sale in 1985.

All species of Kangaroo paw are restricted in distribution to the southern areas of Western Australia. Anigozanthos flavidus occurs in the extreme south west of Western Australia growing from Augusta east to Two Peoples Bay and north to Waroona. It can be found on roadsides, river banks, swamps, shallow water and in eucalypt forests, tolerating light shade. The species A. manglesii is widely distributed throughout the Southwest Australian bio geographic regions, preferring white, yellow or grey sand or sandy loam. The occurrence is confirmed in the northernmost part of its range in the Geraldton Sandplains and the Swan Coastal Plain near Perth. Extensive records also exist of occurrence in Jarrah Forest and Warren region to the south but not reaching the southern coasts. It extends inland to the Avon Wheatbelt.

Lilium is a popular genus of ornamental bulbous plants, belonging to family Liliaceae. Lilium is originated from a Greek word “li” meaning whiteness. It is an herbaceous perennial plant having scaly bulb. Despite the range of forms and their popularity as garden plants, there are two main groups of commercial importance as cut flowers i.e. Asiatic and Oriental Hybrids. They are the result of interspecific hybridization. However, intraspecific and interspecific hybridization face the hurdles like interspecific incompatibility, embryo abortion, degeneration of ovule, poor seed set and pollen sterility. There are several breeding methods and techniques which have been used to overcome these hurdles.

ORIGIN AND DISTRIBUTION

All species of Lilium are native to northern hemisphere. The northern limit extends up to Canada and Siberia, whereas in south, this extends up to Florida and the Nilgiri mountains of India. Majority of lilies known to cultivation are native to the northern hemisphere, chiefly South Canada, Siberia, Florida and India. It is distributed to east coast of Asia, the west coast of North America and the Mediterranean region. Several species i.e. Lilium nepalensis, L. wallichianum and L. polyphyllum are native of Himalayan region and L. niligiriensis have originated from Nilgiri hills. Few species have also been found in north east India. The Far East is the home of almost half of the world lilies and most garden lilies grown in the world have originated there.

The Assyrian monuments dating back to 1000 BC with sculptured lily forms is the first authentic record. The Palestinian coin makers were stamping lily on shekels before 143 BC. Lilium longiflorum popularly known as Easter lily became a florist flowerwhich existed in the Japanese gardens before trade opened the orient to the west. After that it was brought to the England by the Royal Horticultural Society in 1819. Lilies reached to Bermuda from England where it flourished well. Lily depicted an ancient Greek pottery, featured in literature and religions as diverse as pagan myths. Today lilies are enjoying a renaissance of recognition. In Italy and France they are among the most fashionable of flowers. In the USA their popularity has increased tremendously. A number of lily bulbs were imported by the US from the Netherlands for both professional forcing and garden usage. In India, lilies are mostly grown in the hills but Lilium longiflorum do well in the plains, especially under subtropical regions. There are about 7000 registered varieties of lily. The International Lily Register is maintained, under international agreement by the Royal Horticultural Society in London.

Lisianthus is among top ten cut flower crop of the world. In short duration of 20-30 years it become top ranking flower crop of world due to its beautiful flowers which bears beauty of different top ranking old commercial flower crops with variation in flower colours, shape, size with long vase life. It looks like tulips as they began to open and as poppies when fully opened, gaps between the petals appears like daisy, ruffled petals resemble carnations and the most important one is the flower buds and opened double type lisianthus resemble rose buds and open flower respectively and there is no doubt that this is a primary reason for consumer acceptance. Double flowered lisianthus are more demanded in U.S. as cut flower about 80% of lisianthus market whereas, single flowered are more dominant in Japanese and European markets. In Japan over 129 million cut stems were sold in 2001 placing it on number one position in Europe, over 122 million cut stems were sold in 2001 and placing it in top ten ranking and in the United States over 14 million cut stems were sold in 2002, it is not only popular as a cut flower but also as a bedding plant and a pot plant in market (Grueber et al., 1985, Halevy and Kofranek, 1984, Roh and Lawson, 1987, Tjia and Sheehan, 1984, Tija and Sheehan 1986, Skrzypczak et al., 1993 and Harbaugh, 2007), a plant bears 20-30 flowers per plant (Skrzypczak et al., 1993).

Lisianthus (Eustoma) is herbaceous perennial in some cases annual and biennial (Farina, 1989, Halevy and Kofranek, 1984, Wilkins and Grueber, 1983 and Harbaugh, 2007) belongs to family Gentianaceae (Oren-shamir et al., 1999) and genus “Lisianthus”, which is also used as the common name for Eustoma grandiflorum, the plant was commonly known as Lisianthus russellianus when first listed in seed catalogues in the early 1980’s in the United State, later it was given scientific name Eustoma grandiflorum (Raf.) Shinn. (synonyms Eustoma andrewsii; E. russellianum; Lisianthus russellianus) (Bailey and Baily, 1976, Everett, 1981 and Harbaugh, 2007). However, Lisianthus is also known as the Prairie gentian or Texas bluebell (Markham, 1996 and Harbaugh, 2007) and Prairie Rose (Harbaugh, 2007). Eustoma is named after the two Greek words Eu (beautiful, good, well) and stoma (mouth) (Uddin et al., 2013). It is a self compatible (Harbaugh, 2007 and Markham, 1996), diploid crop (Markham, 1996). The buds are colourless is a great disadvantage as an ornamental growth. The range of flower colours available in commercial lines is limited to white and purple, shades of cream, pink, mauve and purple (Markham, 1996 and Harbaugh, 2007).

E. grandiflorum has a set of pharmacologically active compounds and therefore can be a valuable therapeutic material, or can serve as a source of pure chemical compound, apart from its strictly ornamental uses (Skrzypczak et al., 1993).

Tulip cut flower is first among the bulbous ornamentals due to its attractive colours and exquisite flowers. Tulip is a member of Liliaceae family, commonly grown from bulbs. The name tulip is derived from the Persian word “toliban” meaning “turban”, which is an appropriate term to describe the flower shape of certain tulips. The mother bulb dies and is replaced by several daughter bulbs, each of which is an annual unit of growth borne in the axils of the mother bulb scale. Tulips are basically grown for three purposes i.e. bulb production, forcing as cut flower and potted plant and for landscaping (Singh, 2006). The world tulip flower industry is dominated by the Netherlands. Another important tulip producing country is France.

ORIGIN AND DISTRIBUTION

The primary gene centre of the genus Tulipa is located in the Pamir Alai and Tien Shan mountain ranges of central Asia. Diversification in this region, resulting in a distribution from Morocco to Western Europe and eastward to Western China. A second gene centre has been found in the Caucasus (Hoog, 1973). However, Bailey (1963) reported that the tulip is native of Siberia, Turkey, Asia Minor, China, Japan and Mediterranean countries of Europe, the species are particularly abundant in central Asia, in the Bokhara region.

Tulips were first seen in Turkey. The vast majority of species occurs in central and western Asia in the countries such as Turkey, Afghanistan, Iran, Kazakhistan, Turkmenistan, Uzbekistan and Georgia. East of this center of distribution, the range of a couple species extends into northern India and far western China but on limited basis. One species of tulip i.e. Tulipa stellata is found wild in the mid and high hills of Himachal Pradesh and T. aitchisonii is native of Himalaya. There are around a hundred species of Tulipa originated from Turkey to the Pamir and Hindu Kush mountains in Asia, which have been used for breeding purposes. In fact, a small number of species can be found in the wild in Europe and those are all restricted to southern parts, primarily in the Mediterranean region as far west as Portugal. There is not a single species of tulip native to Holland, the country frequently associated with these colourful and well known flowers.

Records show that tulips were in cultivation in Turkey as early as 1000 A.D. In 1578, first tulip appeared in England. Carolus Clusius planted the first tulip in 1593 in the botanical garden in Leiden (The Netherlands). The tulips appear in France in 1598. During 1600-1650 tulip become an exclusive garden plant and it was planted in strategic places in various gardens. Commercial cultivation of tulip started during 1600 in South Haarlem. The famous first parrot tulip was described in 1630. The famous tulip firm namely Voorhelm was established in 1650 in the Haarlem. An inventory of 126 different tulips was released in 1661. The bulbs of tulip had been sold to the U.S. from Holland as early as the eighteenth century. During nineteenth century, several hybrids were developed and cultivable area expanded largely. A fragrant variety Prins van Ooslenrijk and a double early tulip Murillo were developed in 1860. The worldwide popular Darwin tulip group was introduced in 1889. The first classified list of the tulip name including cultivation practices and flower management was released during 1914-1915. The most of the modern tulip complex is based on T. gesneriana with some important hybrids with species tulip. There are approximately 100 of these wild or species of tulips that occur naturally in the world and several thousands of distinctly different hybrids, selection and mutants that are gracing gardens and becoming remunerative to the growers around the world. In India its cultivation is limited to amateur gardeners of hilly areas in Kashmir and Himachal Pradesh. Srinagar, Kangra, Kullu, Mandi, Solan, Shimla, etc. are most suitable areas. Tulip cultivation can be extended in new areas like hills of Uttarakhand in India.

Zantedeschia commonly known as Calla lily, Richardia or Pig lily belongs to family Araceae. It is the only genus in the tribe Zantedeschiae which is the sub family philodendroideae. The name Zantedeschia was given as a tribute to Italian botanist “Giovanni Zantedeschi” (1773-1846) by the German botanist Kurt Sprengel (1766-1833). It is considered by many people in the Republic of Ireland to be a symbol of Irish Republicanism. The Romans valued them so much that they often decorated the edges of the bloom with filaments of gold. These spectacular flowers are beginning to rival the rose in popularity for bridal bouquets. In India there has been initiation in the production of tubers of Calla and few tissue culture labs have started production for New Zealand and Dutch Companies. It is used as potted plants, callas are very attractive. It is used for decoration throughout the spring and early summer months focal point for a spectacular arrangement or bouquet.

ORIGIN AND DISTRIBUTION

The Zantedeschia albomaculata are native to areas of South Central Africa which are subjected to periodic drought (Letty, 1973). Zantedeschia was originally placed in the Calla genus by Carl Linnaeus, Zantedeschia aethiopica was moved in 1818 by Carl (Charles) Sigismund Kunth to the Richardia genus, until it was finally classified as Zantedeschia. Although Zantedeschia did not remain in the Calla genus, even after all this time this wonderful bulb is still referred to by its common name of calla lily.

The name “aethiopica” probably denotes South Africa, the country of the origin. Formerly its name was Calla or Richardia. This plant was introduced in Europe very early, apparently before Van Riebeeck had established the refreshment station at Cape. It is also illustrated in an account of Royal Garden in Paris in 1664. It was sent as one of the interesting plant of Cape to Europe by Simon van der Stal sometime before 1697. The genus is restricted to African continent with seven recognized species Z. aethiopica, Z. albomaculata, Z. ellottiana, Z. jucunda, Z. odoratum, Z. petlandii, andZ. rehmannii.

The common calla (Z. aethiopica) is found from the western Cape through the eastern Cape, Kwa Zullu-Natal, Mpumalanga and in to the Northern Province. In Australia calla lily is naturalized in moist niches of the cooler parts of the New South Wales, southern and south western Victoria, moist valley in Adelaide hills and north west Tasmania. They are serious weeds of pastures and natural bush land in south west of western Australia.

The calla lily is used throughout the world as an ornamental for its flowers. It is mainly grown in temperate and sub tropical climates of the world. They also thrive well under tropical conditions. New Zealand is the topmost producer followed by The Netherland. The original summer flowering calla species were introduced in New Zealand in the early twentieth century. Finland and Scandinavia are also big producer of callas and exported to Sweden and Norway. New Zealand offers a favourable climate to grow callas outdoors and for exports it is grown under controlled conditions. In India it is a potential emerging crop and requires improvement programme for commercialization.

China aster (Callistephus chinensis L. Nees) belongs to family Asteraceae. It is a half hardy and branching annual. Its generic name Callistephus is derived from two Greek words “kalistos” meaning beautiful and “stephos” meaning a crown alluding the large colourful flower heads. Plant is erect having hispid-hairy branches (Bailey, 1963). It is propagated through seed. It is cultivated as cut flower both in the field and under protected condition in temperate countries (Post, 1956 and Warren Auman, 1980). Whereas, in India it is widely grown in Karnataka, Tamil Nadu, West Bengal and Maharastra for cut as well as loose flower production (Janakiram et al., 2001).

ORIGIN AND DISTRIBUTION

China aster is originated from China. Several literatures also stated that China aster is native to China and Japan (Anon., 1950, Bailey, 1963 and Cockshull, 1985). The evolution of China aster has remarkable variations. The present day aster has been developed from single form of wild species, C. chinensis, a plant native to China. The plant was introduced in Europe in 1731 by Jesuit Missionary R.P. d Incarville.

The original plant had single flowers with two or four rows of blue, violet or white ray florets. The first change in the flower type had been the prolongation or development of central florets which lead to quilled flowers. Much original work to improve the plant was done by French. Later the British followed suit with breeding programmes. Most of the recent improvements are the work of American breeders. French florists contributed much in development of double forms. German scientists also evolved many doubled cultivars. These asters were known as German asters because the first great advances in the evolution of the plant were made in Germany and the seeds then used came largely from that country. The comet types introduced in 1886 or 1887 subsequently replaced the quilled types. With the introduction of new branching types by James Vick in 1893, the USA became the main centre of development of this plant as well as for the production of seeds. Since its introduction into Europe, it is grown in almost all the countries of the globe as a garden annual. A good amount of work on China aster on improvement and growing practices has been done in India (Singh, 2006).

Chrysanthemum (Dendranthema spp.) belongs to family Compositae (Asteraceae). It was named by Carolus Linnaeus from the Greek prefix “chrys” means golden and “anthemon” means flower. Chrysanthemum morifolium is the modern florists chrysanthemum, which evolved as a result of endless variation and hybridization from C. sinensis, C. indicum and C. ornatum (Bailey, 1963). This group is also distinguished as C. hortorum. The name of C. morifolium Ramat has been changed to Dendranthema grandiflora Tzvelev (Anderson, 1987, Heywood and Humphries, 1977). Chrysanthemums are mainly herbaceous perennial and bear flowers in capitula. Chrysanthemum perennates in nature by means of stolons and is readily propagated through basal stem cuttings. It ranks second next to roses in importance among the cut flower crops in the world.

ORIGIN AND DISTRIBUTION

Chrysanthemum is originated in northern hemisphere (Europe and Asia). However, some workers reported that China is the native place of chrysanthemum (Carter, 1980). Most of the species in the lineage of present day cultivars are from China.

Chrysanthemum was first cultivated in China as a flowering herb and is described in writing as early as 15^th century B.C. As herb, it was believed to have the power of life. In 18^th century, they appeared in Japan as the crest and official seal of the Emperor. Japan also has a national chrysanthemum day, which is called the Festival of Happiness. In the western world, they are introduced in England, Japan, France and U.S. have developed a wide range of colours, shapes and sizes of flowers. Since the chrysanthemum was first introduced in the U.S. during colonial times, its popularity has grown such that mums now reign as undisputed ‘Queen of the Fall Flowers’. The Netherlands is the foremost importer of chrysanthemum cut flowers and is the centers for re-export to other parts of the world. In India, chrysanthemum is successfully grown under protected as well as open conditions in different states.

Modern rose (Rosa x hybrida) belongs to family Rosaceae. Rose is derived from Greek word “Rhedon” means fragrance. From the ancient time its beauty and attractiveness drag the attention of breeders to create new colours and types. Present cut roses are complex hybrids developed through interspecific hybridization, polyploidy with male and female sterility and these heterozygous genotypes are propagated vegetatively. Rose occupies the first rank among the cut flowers in international market and in India it occupied first position in cut flower export trade.

ORIGIN AND DISTRIBUTION

Roses have been in cultivation since the origins of civilization (Wylie, 1955). The evidence of rose cultivation is found about 5000 years ago in ancient civilization of China, western Asia and northern Africa (Shepherd, 1954). About 120 known wild species, all indigenous to temperate regions of northern hemisphere have been reported to be growing. Eleven species are reported to be growing wild in India. Among the rose species used for perfume extraction purpose Rosa alba originated perhaps in Crimea in Russia, R. borboniana originated in Isle of Reumon in 1817, R. centifolia is native to eastern Caucasus and is under cultivation since 1596. Original home of R. damascena is unknown and it was introduced to Europe from Damascus in Asia Minor during the sixteenth century. R. moschata is native to Middle East and Himalayas and is found at altitudes ranging from 1000 to 4700 meters extending from Europe. The Persian Musk, one of the forms of R. moschata is very common in India. It was introduced into England in 1599. Rosa gallica, known as French rose is native to central and southeast Europe and is in cultivation prior to 1542.

Significant information about roses in antiquity can be seen in the writings of the Greek historian Herodotus (490-420 B.C.), the Greek philosopher Theophrastus (372-287 B.C.) and the Roman naturalist Pliny (23-79 A.D.).

Originally all roses grew wild. About 120 known wild species of roses, all indigenous to the temperate regions of the northern hemisphere, have been reported. The species growing wild in India are Rosa brunonii (Himalayan Musk rose), R. eglanteria, R. gigantea (syn. R. odorata var. gigantean), R. involucrate (syn. R. clinophylla), R. leschenaultiana syn. R. sempervirens, R. longicuspis, R. macrophylla, R. moschata (Musk rose), R. rubiginosa (Sweet Briar, Eglantine rose), R. walpoleana, R. sericea (Ladakh rose). Till the nineteenth century only four species, R. gallica (Red rose), R. canina (Dog rose), R. moschata (Musk rose) and R. phoenicia (Phoenician rose) played a role in the development of cultivars grown at that time.

Tuberose (Polianthes tuberosa L.) belonging to family Amaryllidaceae is half hardy, perennial, bulbous plant. Bulbs are made of scales and leaf bases and stem remains concealed within scales. The generic name Polianthes is derived from Greek word “polios” meaning shining or white and “anthos” meaning flower, in allusion to the blooms of the common tuberose and species tuberosa, the plant being tuberous in nature. Tuberose is commercially cultivated in tropical and sub tropical areas for cut and loose flower trade and also for the extraction of it’s highly valued natural flower oil.

ORIGIN AND DISTRIBUTION

Tuberose is a native of Mexico (Bailey, 1976) from where it spread to different parts of world during 16^th century. The name tuberose is derived from tuberosa, this plant being the tuberous hyacinth as distinguished from the bulbous hyacinth. The name therefore, is tuber-ose, not tuberose. By the middle of the 18^th century the tuberose was still referred for as Hyacinthus indicus tuberosus. Originally Linnaeus had used the name Polianthes floribus alternis in his Hortus Cliffortianus. Later it was grouped under the genus Polianthes and the species tuberosa.

Tuberose is cultivated on large scale in China, Egypt, France, Italy, Israel, South Africa, New Zealand, Spain, Sri Lanka, North Carolina, USA and in many tropical and subtropical countries.

In India tuberose is commonly known as Gulchari and Gulshabbo in Hindi, Rajanigandha in Bengali, Sukandaraji and Nelasampengi in Telegu, Nilasampangi in Tamil and as Sugandharaja in Kannada.

In India its commercial cultivations is confined to Bagnen, Midnapur, Krishnanagar, Ranaghat, Kolaghat and Pariskura in West Bengal, Guwahati and Jorhat in Assam, Pune, Thane, Sangli, Nasik and Ahmednagar in Maharastra, Udaipur, Ajmer and Jaipur in Rajasthan, Navsari and Valsad in Gujrat, Ambala, Gurgaon and Faridabad in Haryana, Ludhiana and Patiala in Punjab, Devanahalli, Tumkur, Kolar, Belgaum and Mysore in Karnataka, East Godavari, Guntur, Chitoor and Krishna district in Andhra Pradesh and Coimbatore and Madurai in Tamil Nadu. Growers of Uttar Pradesh, Delhi and Orissa are also concentrating in tuberose cultivation after evolvement of improved varieties namely Shringar, Suvasini, Prajwal and Vaibhav by Indian breeders (Singh, 2006).

Cyclamen (Cyclamen persicum Mill.), belonging to family Primulaceae, is cultivated throughout the Temperate Zone. Many people have enjoyed the charming plants as potted and/or garden plants. The cyclamen is one of the most famous and important commercial ornamental plants in many countries. It is commonly grown for its flowers, both outdoors and indoors in pots. It is used on a terrace, in flower boxes, in hanging baskets or in beddings. Several species, particularly Cyclamen hederifolium, are hardy and can be grown outdoors in mild climates such as Northwest Europe and the Pacific Northwest of North America. The cyclamen commonly sold by florists is C. persicum, which is frost-tender. Selected cyclamen cultivars can have white, bright pink, red or purple flowers. The use as cut flower is not important part of cyclamen production (Grey-Wilson, 2002), but not rare in Europe, especially in Germany. Some cultivars for cut flower production were bred in Europe, but cut flower cyclamen production is not popular in USA and Japan. Its popularity as pot plant led to many studies on the group, including cytology (Bennett and Grimshaw, 1991), hybridization (Gielly et al., 2001 and Grey-Wilson, 2002) and phenology (Debussche et al., 2004).

ORIGIN AND DISTRIBUTION

Cyclamen originated from extreme eastern Mediterranean region and naturally grows in countries such as Palestine, Syria, Israel, Greece, Turkey, Tunisia or even Mediterranean islands: Rhodes, Cyprus and Sicily (Grey-Wilson, 2002). Cyclamen was introduced in Europe at the end 16^th century, beginning of the 17^th century, passing first through France. Somewhat neglected in the following century, the cyclamen became fashionable again in France in the 1900s. But since 1860, the first variants of the botanical or “wild” cyclamen appeared in England, then in Germany, with much bigger flowers and with varying colour shades.

Freesia is ornamental flowering plant belongs to family Iridaceae subfamily Ixioideae. Freesia is named after a physician and botanist from Germany named “Friedrich H.T. Freese” (1795–1876). Dr. Christian P. Ecklon, a collector of plants from South Africa, named the flower after his friend and student. Modern freesias are known as Freesia x hybrida and cultivated for cut flowers. Freesia is grown for two purposes i.e. cut flower and pot plant.

It is excellent cut flowers because of its appealing shapes and found suitable for various types of arrangements and bouquets. Wide range of colour increases its beauty. Hence, its flower demand increased dramatically in recent year in global market. Forced freesias are grown as pot plant. Colours in freesias consist of pink, red, orange, yellow, blue, lavender, cream and white. Some flowers are multicoloured and possess several variations of colour within the same bud. Because of their delectable fragrance, Freesias have been drafted as a vital ingredient in many fragrance oils. Freesia fragrance oils are utilized for many oil burners and vaporizers. It is used as an essential fragrance in potpourris. Freesia essence oil is high grade essence oil. This is because it is undiluted, alcohol free, and long lasting. The oil is used for aromatherapy to scent candles, soaps, massage oils, bath oils and perfumes.

ORIGIN AND DISTRIBUTION

Freesias are South African in origin. Most species were found in the Cape Province area in southern Africa and two others were found just north of the equator in a tropical climate and even in Sudan. Freesia x hybrida is the name of modern Freesia.

In 1866 Freesia was described as a distinct genus. Freesia alba was first described in 1878 by G.L. Meyer, and became well known in Europe, as F. refracta var. alba. Gumbleton recognized Freesia alba as a distinct species in 1896, but Freesia alba first appeared in the English nursery trade in 1878 and quickly spread to Europe and North America. It appears to have caused quite a sensation, and appeared in almost every horticultural publication of note in both Europe and America in the years following its introduction. Breeding began immediately after F. alba appeared on the market and continues to this day.

Most of the modern hybrids were first made in the 19^th century from a few species. The genus contains some superbly ornamental and adaptable plants. These range from the lovely, scented Freesia alba and the brilliant red Freesia grandiflora and Freesia laxa, to the bizarre green-flowered Freesia viridis.

Gypsophila is herbacaeous annual or perennial flowering plant belongs to family Caryophyllaceae. It contains several ornamental species of which Gypsophila paniculataL. is the only species used as a cut flower (Zvi et al., 2008) and it is applied as filler in flower arrangements (Jones, 1995 and Rehman, 2002). Gypsophila derived from “gypson” meaning chalk and “phylos” meaning lover, an etymology which alludes to its preference for calcareous soils (Vettory et al., 2013). Mainly found in temperate or warm-temperate regions in the Northern Hemisphere, best developed in the Mediterranean region and Near East (Ataslar et al., 2009 and Cronquist, 1981). It is a perennial plant often grown commercially as an annual crop. Root is rich source of triterpine saponins (Henry, 1993, Hostettmann, 1995 and Gevrenova et al., 2010) which are used for commercially in medicines, detergents, adjuvents and cosmetics (Hostettmann, 1995, Tschesche and Wulff, 1973 and Gevrenova et al., 2010). G. paniculata is one of the ten best selling cut flower species worldwide and thus becomes for the genetic improvement and selection of new varieties of commercial interest. Main regions of gypsophila production are Italy, Israel, Belgium, Germany and Holland while currently its cultivation is spreading across to other countries such as Ecuador, Egypt and Kenya due to lower labour costs. Infact, Gypsophila is among one of the most important varieties of cut flowers in Israel, where it is grown in more than 200 hectares of greenhouses and is almost entirely exported. For these reasons, the competition has increased dramatically, making the world market increasingly difficult for companies which feel the need to introduce new varieties that respond better to characteristics desired by consumers and florists.

Stock or Gillyflower (Matthiola spp.) belongs to family Brassicaceae. It is a hardy annual, biennial and perennial plant, flowering in terminal clusters. Matthiola incana is ornamentally important and grown for various purposes i.e. cut flower, fragrance, pot culture, etc. This species develops a woody base in areas where it grows as a biennial or short-lived perennial, hence the name of stock. Generic name “Matthiola” was given in the honour of “Dr. Pietro Andrea Gregoria Matthiole” (1501-1577), personal physician of Emperor Maximilian of Austria. “Incana” means “hairy or gray-white” in reference to plant leaves. It exists as both single and double forms and is valued for fragrant flowers that can be used as fresh or dried cut flower. They are excellent pot plants for the greenhouse in winter and spring and provide good cut flowers. Flowers are well arranged on a long column and colours are variable from white to rose, crimson, purple, yellow, mauve, pink, etc. The highly fragrant flowers are used as a garnish, especially with sweet deserts. Many old names were given to Matthiola incana including stocks, sea stocks, wallflowers and wall or gillyflowers.

ORIGIN AND DISTRIBUTION

Matthiola incana is native to the Iberian Peninsula, France, the Apennine Peninsula, the Balkan Peninsula, the British Isles, Turkey and Cyprus. According to Greuter et al. (1986), four species of Matthiola are known in Sicily and its surroundings: M. fruticulosa(L.) Maire, M. incana (L.) R. Br. & Ait., M. tricuspidata (L.) R. Br. and M. sinuata (L.)R. Br. Unlike its congeners, which only have a single taxon in this archipelago,M. incana consists of three infra-specific subdivisions (Lojacono-Pojero, 1888 and Greuter et al. 1986): ssp. incana (L.) R. Br., which is widespread in other insular and continental Mediterranean enclaves, and the narrow endemics ssp. pulchella (Conti) Greuter & Burdet (confined to the islet of Pantelleria) and ssp. rupestris (Raf.) Nyman, distributed in the surroundings of Palermo (Sicily) and in the Aeolids, the Egadian and Ustica, respectively (Raimondo et al. 1994).

Stock earlier known as Leucoium album or Viola alba was later renamed as Mattiola after Dr. Mattioli, an Italian physician and botanist. This was being grown by the ancient Greeks and was linked for its frangrance. In the wild state, stocks are found in the Mediterranean, Egypt, South Europe and in South Africa and two species, Mattiola incana and M. sinuate are among the rare British natives. Ruth et al. (2009) also reported that Matthiola is widespread in the Mediterranean and Irano-Turanian regions. Both single and double forms were grown in Europe from 10^th century onwards and became popular garden flowers. Apart from its countries of origin, Matthiola incana can be found in Australia, Austria, Bolivia, Bulgaria, China, Colombia, Ecuador, Finland, France, Germany, Greece, Ireland, Israel, Japan, Kazakhstan, Mexico, The Netherlands, New Zealand, Peru, Romania, Sweden, United Kingdom and the United States.

Crossandra belongs to family Acanthaceae. It is also known as Kanakambaram in Tamil or Kanakambara in Kannada and Fire cracker in English because of cracking sound during opening of seed pod. Crossandra is derived from Greek word “krossoi” meaning fringe and “aner” meaning male i.e. fringed anthers. Crossandra is an evergreen shrub and one of the important loose flower used for adornment. The flowers are commonly grown for flower adornment in the form of garlands, venis and gajras. Though it is not fragrant, flowers are very popular because of its attractive bright colour, light weight and good keeping quality. These are used for making garland, either alone or in combination with jasmine flowers. Using crossandra flowers in combination with jasmine is becoming increasingly popular in India, particularly in southern parts, because the jasmine flowers provide colour contrast and the desired fragrance. Crossandra varieties show a remarkable range of colours, varying from orange, pink, red, yellow and double coloured blue-type with white throat.

ORIGIN AND DISTRIBUTION

Crossandra is native to Indo-malaya region. They are all native to sub-tropoical regions including India, Southeast Asia, Africa, and Madagascar.

Crossandra Salisb is a genus of about 50 species which are shrubs or herbs from Tropical Africa, Madagascar and Arabia, where it is grown as a perennial shrub. Crossandra undulaefolia Salisb. (Syn C. infundibuliformis Nees) is a native of East Indies. However, it is sometimes considered to be native of south India and Ceylon. C. guinensis is native of east Africa while C. nilotica has its origin in east Africa and Mozambigue. C. flava is a native of tropical west Africa.

The crossandra plants are distributed in tropical as well as subtropical regions of the world such as south Asia, South America, South Africa and Madagascar. In India, it is commercially cultivated in southern states of Karnataka, Tamil Nadu and Andhra Pradesh (Singh, 2006).

Jasmine (Jasminum spp.) belongs to the family Oleaceae, order Oleales are climbing, trailing or erect shrubby plants and there are both deciduous and evergreen species. The generic name Jasminum is derived from Arabic word “Jassamine”, “Jasmin” and “Jasminum” (Bailey, 1963). Flowers are white, yellow or rarely reddish. Flowers are used for making garlands and Veni. Jamine oil extracted from the flower is highly valued in perfumery industry. France, The Netherlands, Italy, Belgium, Algeria, Turkey, Morocco and Tunisia are major jasmine oil producing countries (Singh, 2006).

ORIGIN AND DISTRIBUTION

Jasmine is native to tropical and subtropical regions; a large number of species are centred on the regions comprising India, China and Malaysia. Jasmines distributed in warm parts of Europe, Asia, Africa and Pacific region, nearly absent from America (Bailey, 1963).

Jasmines have adorned the gardens of central Asia, Afghanistan, Iran, Nepal and many other tropical and sub tropical countries. The distribution patterns of nearly 72 species in India, Malaysia and China provide a strong base to claim that India is one of the principal centers of origin of Jasminum species. Among these, about 40 species are reported to occur in India and mostly cultivated in South India (Bhattacharjee, 1980). Large number of species are however, centered to the region comprising of The Himalayas, China and Malaysia. Other countries where jasmine is cultivated are France, Italy, Morocco, Algeria, North Africa, Spain and Egypt. The cultivated jasmines fall into four species viz., Jasminum sambac, Jasminum auriculatum, Jasminum grandiflorum, and Jasminum multiflorum (Kakada).

In India, the three species Jasminum sambac, Jasminum auriculatum and Jasminum grandiflorum have been mentioned in the ancient Tamil Literature of ‘Sangam’ period (500 B.C. to 200 A.D.). Jasmines are commercially grown in Karnataka, Tamil Nadu and Andhra Pradesh. Other growing states are Madhya Pradesh, West Bengal, Uttar Pradesh and Bihar.

Marigold (Tagetes spp.) an annual flowering plant belongs to family Asteraceae, alternately known as Compositae. The word “marigold” means “Mary’s Gold” and refers to the “Virgin Mary”. However, word “Tagetes” named after the Etruscan God “Tages” who supposedly emerged from the earth as it was being ploughed and was imbued with the power of divination. Cultivated species include Tagetes erecta commonly referred as African marigold and Tagetes patula as French marigold. Marigold commonly propagated by seed and cutting is a leading loose flower of India. Selection of variety is favoured by a higher proportion of double inflorescences.

ORIGIN AND DISTRIBUTION

Origin of marigold is central and South America especially Mexico (Kaplan, 1960). Bailey (1951, 1963 and 1976) reported that marigold is a native to Arizona and Mexico to Argentina.

During early 16^th century it spread to various parts of world from Mexico. African marigold (Tagetes erecta) first reached Spain and from there it become popular as rose of Indies in Southern Europe. Major work on improvement and cultural practices on marigold have been done in USA, Switzerland, France and Germany. In India, marigold was introduced by Portuguese and it became popular and spread quickly because of its wide adaptability to varying climatic and soil conditions. Marigold is well distributed in different states and union territories of India (Singh, 2006).

SPECIES

Earlier, Rydberg (1915) reported that genus Tagetes comprises about 33 species and Bailey (1951 and 1976) mentioned over 30 species, however, Turner and Nesom (1993) reported that Tagetes is made up of nearly 55 species distributed in the American continent, most of them located in Mexico. The two species Tagetes erecta and Tagetes patula are commercially important. Details of these species are given below.

Cattleya belongs to the family Orchidaceae. It is highly important in floriculture and has been cultivated since the 19^th century. The name “Cattleya” was given in the honour of “Sir William Cattley”. Cattleyas have earned the reputation as the “Queen of Orchids” and are known to the public as the ultimate in floral corsages. While no longer the reigning queen of the orchid floral industry it is difficult not to be impressed by a well-flowered Cattleya. No longer limited to white and various shades of lavender and purple, high quality flowers occur in all colours except true blue and black and in a wide range of plant sizes. They are widely known for their large, showy flowers and were used extensively in hybridization for the cut-flower trade until the 1980s when pot plants became more popular.

ORIGIN AND DISTRIBUTION

The genus was named in 1824 by John Lindley after Sir William Cattley who received and was the first to bloom a specimen of Cattleya labiata. William Swainson had discovered the new plant in Pernambuco, Brazil, in 1817 and shipped to the Glasgow Botanic Gardens for identification. After the first importation from Mr. Swainson, no more plants similar to the original Cattleya labiata were discovered for many decades, although many varieties of this species were found. Each variety differed in some way from the plant of Mr. Cattley, either in colour, blooming season or some similar horticultural aspect. Brazil is the centre of distribution, where more than half the species are found (Pabst and Dungs, 1975 and Cruz et al., 2003). They are found naturally only in the western hemisphere, occurring along with Laelias in three major regions of Tropical America i.e. Mexico and Guatemala, northern part of South America from the western Cordilleras of Colombia to British Guiana and the maritime provinces of southern Brazil. Within these three large regions are found most of the species of Cattleya except for C. dowiana in Costa Rica and C. skinneri var. autumnalis from Panama. The plants occur at considerable elevation where the mean temperature is much lower than the mean temperature for the latitude at sea level.

Cymbidium is a genus in the orchid family Orchidaceae. It was first described by Olof Swartz in 1799. The name is derived from the Greek word “kumbos”, meaning “hole”, “cavity”. It refers to the form of the base of the lip. The flowers are long-lasting, both on the plant and when cut large attractive and available in a wide range of colours. Smaller plants are now coming into vogue as decorative pot plants. These have smaller flowers but they are as desirable as the standard varieties and often more acceptable in modern homes. The popularity of miniature cymbidium is now spreading from Asia to the worldwide orchid community. These ancient flowers have been treasured in numerous societies for nearly two thousand years. Many orchid lovers are most charmed by its fragrance and form rather than its petite size.

ORIGIN AND DISTRIBUTION

Cymbidiums figure in the earliest historical records of human association with orchids from China and Japan. Cymbidium native of the monsoonal forest foothills of the Himalayas with the warmer growing Asiatic species from the tropical lowland monsoonal forests. The species come from the Himalayan regions of India and Nepal running through China and Japan, Indo-China down to Malaysia to Australia. This genus is distributed in tropical and subtropical Asia such as northern India, China, Japan, Malaysia, the Philippines, and Borneo and northern Australia. The larger flowered species from which the large flowered hybrids are derived grow at high altitudes. They have been cultivated for thousands of years especially in China. Cymbidiums became popular in Europe during the Victorian era. One feature that makes the plant so popular is the fact that it can survive during cold temperatures as low as 7^oC. Originating in temperate and tropical parts of India, Nepal, China, Japan and down to Malaysia and Australia, cymbidiums are probably the most widely grown of all orchids because of their ease of cultivation and long-lasting flowers. From the 40-50 species, thousands of hybrids have been bred with flowers ranging in colour from white, yellow and orange to pink, red, brown and green, and varying in size from 5-10 cm across. There are several species existing in the Indian forest area. Several species are existing in the forest of Nagaland (Changkija et al., 1992), some of them are given in Table 1.

Dendrobium is a huge genus of orchids. It was established by Olof Swartz in 1799. The name is from the Greek word “Dendron” means “tree” and “bios” means “life”; it means “one who lives on trees”, or essentially, “epiphyte”. The truly spectacular genus Dendrobium contains the largest diversity of horticulturally interesting specimens. More than 1,000 natural species make dendrobium the second-largest orchid genus. Dendrobiums are among the most commonly encountered orchids in the retail trade. Flowers are very long-lasting both on the plant and once cut, looking fresh for up to three weeks or more. Faded flowers on the bottom of the stem can be removed to encourage upper buds to open. Dendrobiums are sturdy orchids that can add a focal point to any room.

ORIGIN AND DISTRIBUTION

Dendrobiums are originally from Southeast Asia. The geographic distribution is from India through China, Japan, Malaysia, Philippines and the islands of the South Pacific with the largest diversity in New Guinea. Dendrobium species are either epiphytic or occasionally lithophytic. They have adapted to a wide variety of habitats from the high altitudes in the Himalayan mountains to lowland tropical forests and even to the dry climate of the Australian desert.

Paphiopedilum is a genus of the orchid subfamily Cypripedioideae and family Orchidaceae. It is derived from Paphos and ancient Greek word “pedilon” means “slipper”. Paphiopedilums are often called “slipper orchids” because of their unique pouch. The pouch traps insects seeking nectar and to leave again they have to climb up past the staminode, behind which they collect or deposit pollinia. Paphiopedilum has a highly ornamental value being the treasure of Orchidaceae. They are easily grown as houseplants and their care is very similar to African Violets.

ORIGIN AND DISTRIBUTION

The genus name Paphiopedilum was established by Ernst Hugo Heinrich Pfitzer in 1886. Ironically, no paphiopedilum occurs on Cyprus- at least not as the genus is understood today. But it was long mixed up with its holarctic relative Cypripedium, which indeed grows in the Mediterranean region. Paphiopedilum was finally decided to be a valid taxon in 1959, but its use has become restricted to eastern Asian species. The genus is native to Indo-Malaysia (South China, Southeast Asia and the Pacific Islands) and India. Paphiopedilum armeniacum is found from China (west Yunnan province) to northern Myanmar. It occurs at 1,600 to 2,000 metres above sea level. It is found growing on limestone hills, in semi-shaded deciduous forest, usually on cliffs above rivers.

SPECIES

Over fifty species of Paphiopedilum have been discovered in the far east from the higher, cooler elevations of the Himalayas, across to China and Southeast Asia and down to the warmer, sea-level regions of Malaysia, Indonesia and the Philippines (Hawkes, 1965). Most species, like Paphiopedilum callosum are terrestrial in nature growing under the shaded and moist conditions of the forest floor (Fitch, 1982). Some species, such as Paphiopedilum niveum are lithophytic found in colonies on the coastal rock formations of the Malaysian islands (Fitch, 1982a). While a few of the larger strap-leaf Paphiopedilum species exposed to the higher light conditions found in their loftier habitats. Paphiopedilum philippinense var. roebelenii and Paphiopedilum lowii provide a strong contrast to such as Paph. lowii, are usually epiphytic Paphiopedilum charlesworthii previously described and serve to illustrate the extent of variation in both plant and flower habit within this genus. Unlike the single-flowered spikes of Paphiopedilum charlesworthii, which are usually around 15 cm tall, those of Paph. philippinense var. roebelenii and Paph. lowii can bear up to a half-dozen flowers and can measure 60 cm or more in length. Paphiopedilums with this kind of flowering habit are referred to as multifloral paphiopedilums (Batchelor). The plants themselves can be quite large particularly in the case of Paphiopedilum lowii, where individual leaves are often well over 30 cm long, though narrow.

Vanda is a genus in the orchid belongs to family Orchidaceae. The name Vanda is derived from the Sanskrit name for the species Vanda tessellate. The name Vandas will be used here to cover all vandaceous orchids including Vanda, Ascocentum, Aerides, Renanthera, Rhynchostylis, etc. and hybrids between them. Although there are some vandaceous orchids that originate in mountainous areas most are low-level plants that love warm temperatures. This has contributed much to the work of hybridists producing flowers for the cut flower market. It is one of the five most horticulturally important orchid genera because it has some of the most magnificent flowers to be found in the entire orchid family. It is highly prized in horticulture for its showy, fragrant, long lasting and intensely colourful flowers.

ORIGIN AND DISTRIBUTION

Vanda was discovered in 1613 by Alvin Semedo, who called it ‘tiao hua’ or ‘air plant’. It is originated from the warm islands of the South Pacific. Places like Malaysia, Thailand, Borneo and the Philippines are natural home to these orchid species. These are mostly epiphytic but sometimes lithophytic or terrestrial orchids. These orchids were distributed in India, Himalaya, S.E. Asia, Indonesia, the Philippines, New Guinea, southern China and northern Australia. About 7 species of Vanda exist in the forest of Nagaland (Changkija et al., 1992). These species are described in Table 1.

Year         Milestones

1951       :    First amino acid was sequenced by F. Sanger and H. Tuppy.

1953    :    J.D. Watson and F.H.C. Crick proposed that DNA is a double helix.

1958    :    Discovery of DNA polymerase by Kornberg’s group and DNA replication studies began.

1959    :    Isolation of RNA polymerase.

1970    :    Discovery of reverse transcriptase showed that RNA can act as the template for the transcription of DNA and opened the way for cloning using cDNA libraries.

1970    :    General method was developed by M. Mandel and A. Higa for introducing DNA into cells by calcium dependent method.

1970    :    G.H. Khorana synthesized an artificial gene from DNA nucleotides.

1971    :    Restriction endonucleases for mapping DNA was first used by K. Danna and D. Nathans.

1972    :    The recombinant DNA Era begins. J. Mertz and R.W. Davis used T4 ligase to join DNA molecules.

Wild relatives of cultivated plants constitute a part of crop genepool, which possess genes that have great potential for their utilization in crop improvement programmes. Wild genepools especially those occurring in biotically disturbed habitats are under threat of genetic erosion and require immediate collection to make use of their wider adaptability, tolerance/resistance to disease, insect-pests, yield, quality attributes and other biotic and abiotic traits. The wild relatives of crop plants occur in all the major phytogeographical regions of India as component of different vegetation types and exhibit variable life forms and habitat specificity. High priority demand is towards collection of those that are at the risk of erosion due to habitat disturbance by manmade/ natural causes and over-exploitation of species for commercial use.

The centres of endemism are also centres of genetic diversity and consequently centres of speciation. Unless suitable conservation methods are applied, important wild relatives of crop plants and rare endemic species in particular have poor chances of survival. Based on the efforts made in the past for collection of diversity in wild relatives of crop plants priorities have been recommended for their conservation on the basis of species status (endemic, rare/threatened/endangered, etc.) and those that have high demand in crop improvement programmes (Arora and Nayar, 1984; Nayar, 1996, 1997; Wiersema and Leon, 1999; Hanelt and Institute of Plant Genetics and Crop Plant Research, 2001).

All economically important species at the risk of genetic erosion require attention for collection and conservation. Highest priority may be assigned to those having highly specialized habitat requirement and those that are endemic/rare/endangered to Indian region. Species producing seeds in large quantity and having easy method of multiplication or conservation are at lower risk than those producing less seed and of the recalcitrant or orthodox types where the storage protocols have been least worked out. Species having short seed viability or low germinability need special investigation.

INTRODUCTION

Heterosis has contributed significantly towards increased crop production and it has become the basis of multibillion dollar agro-businesses in the world (Phillips, 1999). Hybridization is the important way that can create genetic variability and make new options. The F₁ population is obtained from the crossing of two homozygous but genetically dissimilar gametes or individuals (purelines in self pollinated crops or inbred lines in cross pollinated crops) that shows increased or decreased vigour over the better or mid parent value. Heterosis is maximum in F₁, generation which diminishes gradually in later generations. Hence, production of F₁, hybrid seeds of different flower crops is considered as the extreme focused area in floriculture industry.

Production of F₁ hybrids in ornamental crops was carried out even before Shull’s classical theory of hybrid vigour which was reported in 1971 for the first time in plant breeding. Benary Seed Company in Germany released F₁ hybrid in begonia (Begonia semperflorens) in the name of Prima Donna in 1909 (Reimann-Philip, 1983). But that time it did not attract the attention of plant breeders. It was during Second World War in 1942, when flower breeders were attracted towards heterosis breeding and started systematic F₁ hybrids. From 1950 onwards, numerous outstanding hybrids have been developed in many other flower crops in several countries like USA, The Netherlands, Denmark, Germany, United Kingdom and Japan, etc. First hybrid in single geranium was produced in 1960, in other flower crops like antirrhinum, pansy, marigold and zinnia in 1965, ageratum in 1966, geranium double in 1970, dianthus, begonia, impatiens and portulaca in 1976-77, gerbera in 1980 and carnation in 1981. In the present scenario of developed floriculture industry, in many countries F₁, hybrids are available in a number of annual flowers like ageratum, antirrhinum, anemones, calceolaria, dianthus, hollyhock, gazania, marigold, flowering tobacco, pansy, portulaca, stock, wall flower, zinnia, etc.

As emasculation is having the maximum share in cost of hybrid seed production, researchers have found several techniques in different flower crops as the alternative of emasculation and that can help in reducing the cost of hybrid seed production. The list of techniques being utilized commercially in different flower crops (Janakiram, 2004) is as follows:

Since floricultural and ornamental crops are grown for aesthetic purpose and are non-edible there is likely to be less concern in bio safety issues compared to other food crops. Hence there is considerable potential for developing transgenics in ornamental crops. Advances in transgenic technology provide new opportunities for manipulation of the genome. These will have significant impact on expanding and diversifying the gene pool of crop plants, introducing specific genes and shortening the time required for the production of new varieties or hybrids. Molecular breeding is beneficial to increase the production and quality by creating plants with enhanced resistance to diseases, insects or viruses and increased tolerance to environmental stresses like salinity, temperature or drought. Through this technique genes for shelf life, flower colour and architecture may be directly transferred to develop new varieties that are tailor made to customer preferences. Bioluminescent orchids and blue roses are the significant outcomes of the genetic transformation studies in ornamentals (Swarnapiria, 2009).

GENETIC ENGINEERING VS. TRADITIONAL BREEDING

Breeding with transgenic plants is only justified when genetic variation within the primary and secondary gene pools of a species is too small or gene transfer by conventional techniques is difficult and time-consuming. There are a number of different means for increasing or even creating new genetic variation, like species hybridization, mutation induction and protoplast fusion, which have been frequently used to breed new varieties. It is worth mentioning that neither plant cultivation nor commercialization of commodities has any legal requirements and a public discussion about these plants is also lacking. Therefore, a breeder must carefully assess the different constraints of transgenic breeding before starting a breeding program which are the legal aspects and acceptance by farmers, consumers and stakeholders (Jung, 2010).

INTRODUCTION

The seed (or zygotic seed) is the vehicle that connects one generation to another in much of the plant kingdom. By means of seed, plants are able to transmit their genetic constitution in generations and therefore seeds are the most appropriate means of propagation, storage and dispersal. A synthetic seed or ‘synseed’ is “an encapsulated single somatic embryo”, i.e., a clonal product that can be handled and used as real seed for transport, storage and sowing and that, therefore, would eventually grow either in vitro or ex vitro, into a plantlet through a process of “conversion”. Now a day, artificial seed technology is one of the most important tools to breeders and scientists of plant tissue culture.

In general, synthetic seeds are defined as artificially encapsulated somatic embryos, shoot tips, axillary buds or any other meristematic tissue, used for sowing as a seeds and posses the ability to convert into whole plant under in vitro and in vivo conditions and keep its potential also after storage (Capuano et al., 1998). The aim and scope for switching towards artificial seed technology was for the fact that the cost-effective mass propagation of elite plant genotypes will be promoted. There would also be a channel for new transgenic plants produced through biotechnological techniques to be transferred directly to the greenhouse or field. The artificial seed technology has been applied to a number of plant species belonging to angiosperms.

Artificial seeds have great potential for large scale production of plants at low cost as an alternative to true seeds (Roy and Mandal, 2008). An artificial seed is often described as a novel analogue to true seed consisting of a somatic embryo surrounded by an artificial coat which is atmost equivalent to an immature zygotic embryo, possibly at post-heart stage or early cotyledonary stage. The artificial seed production technique was first used in clonal propagation to cultivate somatic embryos placed into an artificial endosperm and constrained by an artificial seed coat. Today artificial seeds represent capsules with a gel envelope, which contain not only somatic embryos but also axillary and apical buds or stem and root segments (Vdovitchenko et al., 2011). Explants such as shoot tips, axillary buds and somatic embryos are encapsulated in cryoprotectant material like hydrogel, alginate gel, ethylene glycol, dimethylsulfoxide (DMSO) and others that can be developed into a plant. The coating protects the explants from mechanical damage during handling and allows germination and conversion to occur without inducing undesirable variations. They behave like true seeds and sprout into seedlings under suitable conditions.

INTRODUCTION

Seed is one of the most vital and important inputs for increasing agricultural production. Commercial flower seed production is an international business involving highly specialized growers. Compared with the production of seeds of cereals, pulses oilseed even vegetable seeds, it is done on a relatively small scale. However, a large number of species are being produced under very different cultural conditions. It seems to have great scope for expansion in developing countries like India, especially under northern India climatic zones having favourable growing conditions, skilled and cheap labour.

Annual flowers like petunia, coreopsis, helichrysum, phlox, nasturtium, marigold, gaillardia, salvia, ice plant, verbena, nemesia, pansy, poppy, larkspur, chrysanthemum, etc. have been widely used for seed production. After release of a variety its seed have to be multiplied in sufficient quantity which takes 3-4 generations before it reaches to the farmers for commercial use. The Indian seed programme largely adheres to the limited generations system for seed multiplication in a phased manner. The system recognizes three generations namely breeder, foundation and certified seeds and during multiplication cycle care has to be taken so that variety does not degenerate but maintains its original characteristics.

Seed production of flower has undergone unrestricted revolutionary growth in term of number of crops and their varieties as well as production. This phenomenal development is the result of technological advancement in the production of flower seed and development in the area of production, packaging, handling, transportation and marketing of flower and ornamental plants all over the world. The world germplasm contains more than 1500 different varieties of flowers and ornamental crops. Each species of flower grown for seed has its own specific time, culture, problem of pollination and harvesting and storage requirements. Mild climate with moderate rainfall favour the growth and development of most flower and ornamental crops.

Production of seeds of annual flower is having immense impetus as regards the diversification in floribusiness. The flower seed industry is truly international, operating in many countries in the world. Large quantities of annual flower seeds are required both by developed and developing countries adding to the beauty of private and large gardens and also for cut flower production. Annual flower are indispensable in respect of beautification of gardens. In the world, many nurseries are engaged in the production of seed of seasonal flowers and are having a blooming business. The great diversity of flower seed crops is fascinating facet of the world seed industry and they are of high value and low volume in contrast to the high volume and low value seeds. The varied agroclimatic condition existing in India provides a vast scope for the production of a wide range of flower seeds of both tropical and temperate origin. With the availability of relatively inexpensive human labour, technical expertise, and other material, it is possible to organize a strong flower seed production programmed exclusively for export purpose.