eChapters

INTRODUCTION

Control of weeds below economic threshold level in potato crop is a substantial component of crop management strategy for achieving optimum productivity. In our country, potatoes are raised eitherin assured irrigation or as rainfedcrop with comparatively heavy doses of fertilizers. Therefore, weedsarebound to become menaceduringactive cropgrowth period of this cropand theseundesired plants needs to be controlled in early stages of crop growth by adopting several techniques. Current scenario of crop production is different from past due to manyreasons. Farm holdings are becoming smaller day-by-day and majority of potato growers are small and marginal farmers. They have limitedcapacity for purchase of inputs and farm machines. They need economical, efficient and easier to adopt weed control technologies.Farm labour is also costly nowin many regions as they are migrating towards cities and industrial townships. This scenariobecomes more complex as weeds would be major challenge due to global warming, which is slowly changingclimatesof potato growingregions in India.Theconcern forenvironmentalissuesemphasizeuponadoptingsystems and integrated approach for managing weeds in such away that thesedo not reducethe potato and system productivity (CPRI, 2014).

INTRODUCTION

Agricultural biodiversity is the basis ofglobal food security. Wide variety of crops, soil and cropping systems are nurtured by human culturaldiversity since times immemorial. Potatois a majorfood crop which is widely acknowledged now. It is consumed by the world’s largest population than any othervegetable, produces more calories and protein per unit of land and time than any other food crop.Potatohas much potentialfor sustainable, non-chemical farming, fits well with many crop rotations,and does very wellwith naturalfertilizers. Over the years chemical based farming has started experiencing reduced production and increased costs, orboth (Singh et al.,2011 and Sreenivasa et al., 2010). Monoculture of crops such as rice, wheat and cotton etc., depletes topsoil of nutrients and reduces diversity of beneficial microbes leading to reduced soil productivity. It is finally making the crop plants vulnerable to parasites and pathogens. Environmental pollution caused bychemical fertilizers and pesticides is posinga serious threat worldwide (Doranet al., 1996). Healthy soil is the foundation upon which sustainable agriculture is built. Farming practices differ mainly based on soil inputs and crop protection measures (Devarinti,2016).Thusthere is growinginterestinorganicallyproducedproducts and farmers are shifting from conventional to organic farming. In India, Sikkim is the firststate where completelyorganic farmingis adopted. Farming practices differ mainly based on soil inputs and crop protection measures (Devarinti, 2016) in different regions. Different types of farming based on use of natural products are described in the chapter.

INTRODUCTION

Potato (Solanum tuberosum L.) is one of the major food crops whichrecorded total globalproduction of 388.1million metrictonnes(mt) in 2017-18. Out of the major staple food crops, potato production (388.1mt) is exceeded only by maize (1077.98 mt), wheat (761.88 mt) and rice (494.88 mt) (FAOSTAT, 2017). According to the three years (2012-2014)averages,globally, India ranks 3^rdin area and 2^ndin terms of production nextto China. Thenutritional quality, high productivity and acquiescence for inclusion in intensive cropping systems of this shortduration crop reflects its great potentialin modern agriculture to feed theexponentiallyrisingpopulationin the developingcountries. But, theintensive cultivation of potato crops urges increased use of fertilizers, pesticides and other chemicals leading to high input costs with plateauingyields. Blanket dose of fertilizers as well as imbalance use of nutrientsnot only increases the cost of farm inputs but also degrades soil condition and causes severe environment pollution. Indiscriminate use of insecticide as well as fungicide is verycommon in potato crop which contaminate the environment and deteriorate product quality. Therefore, the chemical inputs need to be optimised based on actual requirement of the crops for sustainable crop production. Furthermore, potato production is associated witha high tillagepractices, number of tillage operation would depend on soiltype, previous crop etc.

INTRODUCTION

Potato is a highly versatile crop grown in varying environmental conditions viz. plains, hills and plateau in India. It is grown under short days (SD) in the plains in autumn season and under long days (LD) during summers in hills. Therefore, the pattern of growth, duration of crop as well as the yield varies according to the environment. Besides, the potato cultivars differ from each other in relation to their morphological characters, response to the environmental factors and finally in yield and the dry matter accumulation. The biochemical reasons for such variations are not clear, but it seems that genetically determined differences in the synthesis of growth regulators such as auxins, cytokinins, gibberellic acid (GA) and abscissic acid (ABA) play an important role and their proportion at different growth stages affect the growth and development and thus, the yield of the cultivar (Minhas and Singh, 2003).The present chapter deals with the physiologicalfactors, which determine the growth, development and yield of potato crop.

INTRODUCTION

Potato is a vegetatively propagated crop mainly through tubers. Seed is the costliest input, which constitutes nearly half of the production cost. Due to the high cost of seed, seed replacement rate is low, which results in build up of various tuberborne diseases.To maintain high tuber yield, multiplication is done through “Seed Plot Technique” (SPT), which involves seed production in low aphid pressure period to minimize transmission of viruses and also adopt a set of cultural practices aimed at minimizing disease spread. The production of healthy seed tubers through SPT is expensive and the low rate of tuber multiplication (normally 68 times) provides only a limited quantity of quality tubers. Further, the low aphid areas suitable for producing healthy seed in the country lie in the northern plains and high hills (> 1800 m above mean sea level). Changing climate scenario is likely to shrink the window for potato cultivation and increase pathogen/vector attacks across the country thereby seriously limiting quantity and quality of seed in the country. An alternative technology of true potato seed (TPS) or use of botanical seed has shown great potential for producing both diseasefree healthy (quality) and cheaper planting materials for commercial potato production. This chapter describes the classical TPS technology and two novel potential TPSbased technologies in potato propagation.

INTRODUCTION

Potato (Solanum tuberosum L.) is the most importantnon-grain food crop of the world and is central to global food security, offeringhigher yields in calories per acre thanany grain.More than 2 billion people worldwide are estimated to be dependent on potato for food, feed or income. Global potato breeding concentrates on the objectives of increasing the yields through resistance to various biotic and abiotic stresses along with improving quality traits but the cultivated potato is an auto-tetraploid (2n=4x=48), clonally propagated, highly heterozygous, and suffers acute inbreeding depression. All these characters complicate genetics/genomic studies as well as breeding efforts to improve important traits such as disease/pest resistance, processing quality and nutritive value.This has lead to the need of high quality, well annotated genome sequence of potato along with high resolution mapsfor aiding the identification of allelic variants for important agronomic traits in potato. Multiple linkage maps have beenconstructedfor potato usingvarious molecularmarkers like RFLPs, AFLPs and SSRs in an effort to develop markers for marker assisted breeding and facilitate map based cloning. These earlierefforts got unequivocal boost when an internationalcollaborationof13countriesviz.Argentina, Brazil,China, Chile, India, Ireland, The Netherlands,New Zealand, Peru, Poland, Russia, the United Kingdom and the United States came together to form the Potato Genome Sequencing Consortium (PGSC) with aim to decipher the potato genome (Xu et al., 2011). The potato genome sequence has provided a major boost to gaining a better understanding of potato trait biology, underpinning future breeding efforts. This would greatly help in enhancing the breeding techniques by implementing the genome assisted methods for imparting the greatertolerance to various biotic and abiotic stressesand ultimately increasing the yields. The knowledge of the complete genome sequence is an invaluable resource for the identificationof genes and variant/novel alleles of genes for everytrait ofinterest to potato breeders. This is very much required for feeding the ever burgeoning global population.

INTRODUCTION

Potato (Solanum tuberosum L.) is 3^rd most important noncereal food crop in the world, after rice and wheat, and popularly known as ‘the king of vegetables’. Potatoes are considered as a nonfattening, nutritious and wholesome food, which supply important nutrients to the human diet. Tubers contain significant concentrations of vitamin C and essential amino acids. Potatoes can be consumed in many ways, including baking, boiling, roasting, frying, steaming, and microwaving. It is integral part of food and traditional cousins and likely to find more importance in the dietary habit. Potatoes are either consumed directly or they are processed to give products such as chips and French fries, mashed and canned potatoes. Besides being important in human diet, potatoes are also used as animal feed and as raw material for starch and alcohol production. Potato was an occasional dish of the affluent class at the time of independence but now it is the most abundantly consumed horticulture based food by the poorest of the poor in the country. At present about 68% are utilized for table purpose, 7.5% for processing; 8.5% for seed and remaining 16% produce goes waste due to pre and postharvest handling.

INTRODUCTION

India has emerged as second highest producer of potato in the world, after China. The crop has been identified as the most important crop for ensuring food security for the developing countries, and a highly nutritious food providing more calories, vitamins and nutrients per unit area of cropped land than any other staple food. The crop has witnessed phenomenal increase over the seven decades of potato research in India. For a crop which was introduced in the country only in the early 17^th century, and was neither adapted, nor its staple food, the remarkable increase in potato production and productivity is an exceptional achievement of breeding this temperate introduction into the present day potato cultivars.

BREEDING REQUIREMENTS

The table potatoes introduced in India were adapted to long photoperiods of up to 14 hours and crop durations of 140-180 days allowing them to be cultivated only in the hilly regions of India. The unique and diverse agro-climatic condition of the country, comprising of contrasting conditions in the Indian plains, where potato could be grown in the winter months necessitated breeding efforts to develop the potato varieties suitable for the sub-tropical plains and the other potato growing regions. The winters in sub-tropical plains have short photoperiod (with about 10-11 hours sunshine) and the crop duration is limited to 90-100 days because of short and mild winter. Further constraints on photosynthetic activity due to early morning fog, posing a strong limitation on tuber bulking. This is followed immediately by a long hot summer period, which coincides with the post-harvest storage period of potato, creating challenging storage problems. The rapid degeneration of seed stocks, with concomitant decrease in yield due to accumulation of viral diseases and tuber storage and utilization in hot and humid climate thus needed to be addressed, in priority. Besides, these southern, north Bengal and Sikkim hills and the plateau offer agro climatic area specific problems related to the biotic and abiotic stresses prevalent in the regions. Moreover, the unique reproductive biology of the plant made it possible to flower only under the long day conditions of the hills, where hybridization was initiated at Kufri (Shimla), Himachal Pradesh. However, during the initial years, it was not possible to evaluate the breeding progenies in the warmer sub-tropical plains due to the quick degeneration of hill bred progenies and dormancy of hill potatoes.

INTRODUCTION

Potato belongs to a genetically diverse genus Solanum having nearly 235 tuber bearing species. These speciesform a polyploid series ranging from diploids (2n=2x=24)tohexaploids (2n=2x=72), with12 asthebasic chromosome number. There is also arichdiversity within the cultivatedspecies of potato and they were initially grouped under six different species. Later studies, however, recognized those species as land race populations of S. tuberosum, with the eight groups: Ajanhuiri, Andigenum, Chaucha, Chilotanum, Curtilobum, Juzepczukii,Phurejaand Stenotomum(Spooner & Salas, 2006). Thisrich genetic diversity of the species is available in the Andean mountain where potato originated. In India,however, it was quite natural that genetic variability within the locally available material will be meager since, it was introduced here only about 400 years ago. On the contrary, more than 500 differentvarieties existed in India by 1940. This paradoxical situation was primarily due to assigning different local names toasingle variety in different parts ofthe country. The available local varieties in India consisted of both early introductions belonging to either pure Solanum tuberosum ssp. andigenum or their hybrids with S. tuberosumssp.tuberosumas well as later European introductions belonging to S. tuberosum ssp. tuberosum.

INTRODUCTION

Potatois an important crop for food and nutritional security of India. Itis mainly grown as rabi crop and 90 per cent of itscultivation is confined to northern Indo-Gangetic plains comprisingthestatesof UttarPradesh, WestBengal,Bihar, Punjab and Haryana. Due to climate change, the main potato season is likely to be warmer than the rest of the seasonsof the year and will affect seriously the potato production in India. Climate change is defined by Intergovernmental Panel on Climate Change (IPCC) as a plausible future climate thathas been constructedfor explicitusein investigatingthepotential consequences of climate change. It is the difference in climate over a period of time (with respect toa base line or a reference period) and corresponds to a statistically significant trend of meanclimate or itsvariability persistent over along period of time. Climate change may be due to both natural aswell asanthropogenic factors. Reference periods are typically of three decadesused as climatological baseline period in impacts and adaptation assessments to Global climate model. The future climate minus baseline climate are used for model scenario comparison of most climate variables. Typically a number of fixed time horizons in the future are produced from model output.For example 2020 (2010-2039), 2050 (2040-2069), 2080 (2070-2099). Future climates are estimated through models, called global circulation models, or global climate models (GCM). GCM simulations are carried out by means of a large set of heavy computations. The IPCC TAR reported 7 GCMs for developing climate scenarios while IPCC FAR reported 21 different GCMs.

Introduction

At the time of inception of CPRI, in the year 1949, India used to produce 1.54 million tonnes potatoes from 0.234 million ha area at an average productivity level of 6.58 tonne/ ha. As per the first advance estimate of the NHB, the potato production in India during 2018-19 was 52.5 million tonnes from 2.18 million ha area at 24 tonnes/ ha productivity. The potato production, area and productivity increased over 7 decades by 34, 9.3 and 3.7 times, respectively. ICAR-CPRI has been adequately recognised by the nation on several occasions for this stupendous contribution. However, potato in India still has to transform from simply a vegetable supplement to serious food security option. Ability of potato to produce highest nutrition and dry matter on per unit area and time basis, among major food crops, made FAO to declare it the crop to address future global food security and poverty alleviation during 2008. Rising number of working couples, rapid rate of urbanization, enhanced tendency of eating out of home, higher disposable income levels of people and important place of potato in fast food items, create an ideal situation for expansion of potato consumption in the near and distant future.

A perusal of various R&D efforts and outcomes in the field of agriculture in general and, more specifically the potato, reveals that “Business as Usual” scenario would not hold much longer. We have to anticipate and get ready to tackle much more complex and diverse future challenges in our respective fields. As the time lag between research and development efforts and the final application/ adoption of the output may stretch over decades, a long term vision and blueprint of action plan is highly important. This chapter attempts to envisage long term state of affairs of potato industry in India and formulate a strategy to fulfil national needs through a well-documented plan to tackle anticipated challenges.

Greenhouse technology development has made good progress in India during the last two decades, and steps necessary for promoting greenhouse cultivation of flower and vegetable crops have been started. Energy-efficient greenhouse cultivation continues to be an area of active research and development, and this is sought to be achieved through precision equipment and protocols. While the gap between the demand and supply of most horticultural crops remains wide and the country plans to double the production of horticultural crops by 2012, the protected cultivation technology holds the key to meet the targets. It clearly emerges that greenhouse technology has multipurpose application for sustainable development of a hilly zone. During harsh winter which otherwise threatens the survival of human beings in these remote hilly areas, popularization and further improvements in greenhouse management could provide a sign of relief in solitude of white snow cover.

The popularity of greenhouse-grown produce continues to rise as it provides a variety of produce during the “off” season. Greenhouses are good for healthy plant nursery production and other a number of crops for profit. Though the greenhouse farming in India like in other countries is progressing but there are certain problems and constraints as stated below.

1.Greenhouse production generally is highly technical and requires constant and continuous attention during cropping period. Relevant information about the crop environmental requirement such as right temperature, humidity etc is essential.

2.Knowledge about the right type of inputs for greenhouse production is a must which is not easily available.

3.Greenhouses cultivation increases the land, water and fertilizer productivity along with reduction in pesticide use. But such interventions are associated with very high initial cost. Although, it is linked with several Government Schemes in India and have 50-90% of subsidies depending on intervention, status / socio-economic condition and policy of State and Central Government.

4.Uninterrupted and regular power supply is required for operating cooling and heating system of the controlled environment greenhouses which necessitate a back-up power pack which is also expensive and underutilized.

Nutrient film technique popularly known as NFT is one of the methods of soilless farming or hydroponics where in plants have their roots in a shallow stream of recirculating water containing all the nutrients required for plant growth dissolved without any solid growing medium. The lower portion of roots remain submerged in flowing nutrient medium in a water tight gully having enough air for the roots. The nutrient film technique was developed during the late 1960s by Dr Allan Cooper at the Glasshouse Crop Research Institute, England. The method is already used in commercial horticulture in many countries including India. NFT is potential for other crops such as animal fodder and even energy-source plants for fuel production.

NFT related technologies

Hydroponics : Cultivation of plants in nutrient solutions is known as hydroponic cutlivation. This ia gerally done under protected structures and in containers arranged horizontally or vrtically (vertical garden). Use of nutrient film or fertilizers is made to feed the plants. Hydroponics can be practiced following nutrient film technique method, trough method, deep water floating method and coco peat bag culture.

Soilless media or culture

Gravel culture is a general term which applies to the growing of plants without soil in an inert medium into which nutrient solutions are usually pumped automatically at regular intervals. Haydite (shale and clay fused at high temperatures), soft- or hard-coal cinders, limestone chips, calcareous gravel, silica gravel, crushed granite and other inert and slowly decomposing materials are included in the term “gravel”. The more important greenhouse flowering crops include roses, carnations, chrysanthemums, gardenias, snapdragons, lilies, asters, pansies, annual chrysanthemums, dahlias, bachelor buttons and others.

Media ingredients and Mix

Commercially available materials like peat, sphagnum moss, vermiculite, perlite and locally available materials like sand, red soil, common manure/ compost and rice husk can be used in different proportions to grow greenhouse crops. These ingredients should be of high quality to prepare a good mix. They should be free from undesirable toxic elements like nickel, chromium, cadmium, lead etc. Most common medium other than soil used in India in greenhouses is coco peat. In other countries these are Rockwool and Peat-vermiculite.

Pasteurization: The heat treatment given to soil to free it of harmful microorganisms or pathogens and pests is called pasteurization. It kills weed seeds also.

Pasteurization of greenhouse plant growing media

Greenhouse growing medium may contain harmful pests such as disease causing organisms, nematodes, insects and weed seeds. It is advisable to decontaminate it by heat treatment or by treating with volatile permissible chemicals.

Flooding

Flooding affects both above and below ground ecosystem processes in greenhouses. Below-ground changes may be less obvious, they are as important as the above-ground changes. Soil microorganisms are sensitive to disturbance, and shifts in soil microbial community structure are expected when anaerobic conditions develop from flooding. The useful aerobic microbes are destroyed by flooding. It takes time to built-up their population in the soil to benefit plants. Due to anaerobic condition created by flooding in greenhouse plant roots in soil are not able to breath, function and hence wilting of plants occurs. Flood irrigation in excess create flood like condition in greenhouses at time. Excess water in soils is a major factor affecting plant survival and functioning. In saturated soils, the supply of atmospheric oxygen into the soil is curtailed and various facultative and obligate anaerobic microorganisms use oxidized compounds as electron acceptors for respiration, thus converting them to reduced forms. The reduction and the associated processes influence plant survival, growth and functioning in wetlands. Flood irrigation should always be just wet the soil not flood inviting stagnation of water.

A chain of reactions is initiated upon soil flooding leading to reduced soil redox potential conditions. These reactions include physical, chemical and biological processes that have significant implications on plants growth. Physical processes include restriction of atmospheric gas diffusion in the soil leading to depletion of soil oxygen and accumulation of carbon dioxide. Shortly after flooding, the limited supply of oxygen in soil pore spaces is depleted rapidly by roots, microorganisms, and soil reductants. This process leads to oxygen depletion and reduction in soil oxidation reduction potential followed by a chain of soil chemical changes. The processes that follow include denitrification, reduction of iron, manganese and sulphate, and changing soil pH

Drainage is the natural or artificial removal of excess water from a surface or sub-surface from an area or field or soil or greenhouse. The internal drainage of most agricultural soils in greenhouse is good enough to prevent severe waterlogging. Water logging creates anaerobic conditions that harm root growth in absence of oxygen. Soils need artificial drainage to improve production or to manage water supplies in certain conditions. Though the roots need moisture but they need oxygen too and poor drainage is g result in roots rotting and death of plants.

The irrigation water inside the greenhouse may accumulate and cause anaerobic condition. Similarly plants in pots should not accumulate water. Water should not stand in pots and beds for longer period. If it stays longer drainage is poor and needs to be corrected. It is better to have good drainage system in greenhouses or protected structures.

Besides irrigation rains water seeps in the greenhouse. The drainage system prevents running or its accumulation in low lying portion of the greenhouse. Drainage system in greenhouses is a must for avoiding damage to the crops.

The following various types of irrigation methods and their systems are used in crop production in greenhouses and plant nurseries.

1.Hand Watering

2.Perimeter Watering System

3.Tube Watering System

4.Drip Irrigation System

5.Overhead Spray System

6.Ebb and Flow System.

About 80% of the world population live in developing countries and are dependent on agriculture. Agricultural products, just after harvesting, are dried to the safe moisture level. Open solar drying for drying agricultural products is being practiced since ages throughout the world. Due to its several drawbacks as danger of (dust, rain, ultra-violet rays, birds, animals etc.), advance technique, i.e. greenhouse drying, is being adopted for drying crops to reduce the drying time and increase the quality of the food products.

Drying involves a heat and mass transfer phenomenon in which heat energy supplied to the product surface is utilized in two ways: (i) to increase the product surface temperature in the form of sensible heat and (ii) to vaporize the moisture present in product through the provision of the latent heat of vaporization. The removal of moisture from the interior of the product takes place due to induced vapour pressure difference between the product and surrounding medium. The moisture from the interior diffuses to the product surface to replenish the evaporated surrounding moisture.

The working principle of greenhouse drying is in which the product is placed in trays receiving the solar radiations through the plastic cover and moisture is removed by natural convection or forced convection.

Greenhouse drying are being adopted to reduce the losses and to increase the quality of the dried products. Solar energy is the most promising power source of energy for drying of agricultural products. Most of the farmers adopt open sun drying because it is abundant and inexpensive. But the open sun dried products are also subjected to remarkable losses due to environment. Losses of fruits and vegetables are estimated to be 30%-40% during drying under open sunlight. The post-harvest losses can be reduced by adopting advanced means of drying, i.e. greenhouse drying. The greenhouse dryer can be operated in natural and forced mode as required. Studies reveal that the greenhouse dried products are of superior quality and better colour retention as compared to open sun drying.