eChapters

Geranium (Pelargonium spp.) belongs to the family Geraniaceae. The name Pelargonium was introduced by Johannes Burman a Dutch Botanist in 1738, from the Greek word “Pelargos” means stork because the seed head looks like a stork’s beak. Due to their long and beak-shaped fruits, they are also known by the name of cranesbills. Geraniums are popular flowering plants and grown as decorative pot plants for indoor and outdoor. There is a world demand for the rose geranium essential oil of Pelargonium species which contains geraniol, linalool and citronellol, substitutes for the expensive attar of roses in the perfume trade. Pelargonium species was known for their medicinal value by the earliest indigenous people (Swanepoel, 2009).

ORIGIN AND DISTRIBUTION

The plants originated from South Africa (Swanepoel, 2009), as well as Reunion, Madagascar, Egypt and Morocco and were introduced to European countries such as Italy, Spain and France in the 17^th century. Geranium includes around 422 species of flowering annual, biennial and perennial plants. Both Geranium and Pelargonium are genera in the Family Geraniaceae. Linnaeus originally included all the species in one genus. Geranium, but they were later separated into two genera by Charles L’ Heritier in 1789. Although members of the Pelargonium genus are often referred to as ‘Pelargoniums’ to try to avoid confusion, many people still use the older common name of ‘Geranium’.

Most of the Pelargonium plants cultivated in Europe and North America have their origins in South Africa. Geranium robertianum is found throughout Europe, Asia and North Africa where it grows on a variety of soils, rocks, tree trunks and decaying organic matter such as logs. It is a component of virtually all forest types there. In the Pacific Northwest it is primarily found west of the Cascade crest although it extends along the Columbia River in Klickitat County. Ecological structure of Geranium robertianum L. populations under natural conditions and in the garden in Poland. Geranium farreri found in China, in south Gansu and north-west Sichuan.

Geranium (Pelargonium spp.) belongs to the family Geraniaceae. The name Pelargonium was introduced by Johannes Burman a Dutch Botanist in 1738, from the Greek word “Pelargos” means stork because the seed head looks like a stork’s beak.  Due to their long and beak-shaped fruits, they are also known by the name of cranesbills. Geraniums are popular flowering plants and grown as decorative pot plants for indoor and outdoor. There is a world demand for the rose geranium essential oil of Pelargonium species which contains geraniol, linalool and citronellol, substitutes for the expensive attar of roses in the perfume trade. Pelargonium species was known for their medicinal value by the earliest indigenous people (Swanepoel, 2009).

Gerbera (Gerbera jamesonii), a tender, stemless, perennial flowering plant, belongs to family Asteraceae. Gerbera was named in honour of the German botanist and naturalist “Traugott Gerber” who was a friend of Carolus Linnaeus. Gerbera is very popular and widely used as cut flower or as a decorative garden plant. It was discovered in 1878 near Barberton, eastern Transvaal (South Africa), therefore, referred as the Barberton daisy or Transvaal daisy. It is ideal for beds, borders and pots and is the fifth most important cut flower in the world.

ORIGIN AND DISTRIBUTION

Gerbera is originated from South Africa. The native distribution of genus Gerbera comprising 30 species, extends to Africa, Madagascar, tropical Asia and South America (Hansen, 1985 and Bremer, 1994). Whereas, it was also reported that gerbera is found in the temperate and tropical parts of Asia and Africa and about 7 species are recorded in India. This plant is distributed in the temperate Himalayas from Kashmir to Nepal. Gerbera is commonly seen on the dry slopes near Shimla and on hillsides above Gulmarg in Kashmir (Anon., 1956). The G. jamesonii is found naturally in grassland in sandy soils in Mpumalanga and Barberton district. The genus Gerbera has approximately 14 southern African species, some of which are found only in Western Cape. G. aurantica is endemic to the mist belt grassland region of KwaZulu-Natal and is known from approximately 9 scattered populations from Helahela in the south to Babanango in the north. This species is under threat of extinction due to habitat fragmentation and degradation. In India they are distributed in temperate Himalayas at altitudes of 1300-3200 metres and observed from Kashmir to Nepal.

The genus Gerbera is named in honour of German naturalist Traugott Gerber who travelled in Russia in 1743 and started first botanical garden in Moscow. However, for species i.e. jamesonii, it is a matter of different opinion. The jamesonii was named for Dr. L. S. Jameson, a British colonial statesman in South Africa who was one of the discoverers of this species, whereas in some literature, the species was named after Robert Jameson who collected live specimens while on a prospecting expedition to the Barberton district in 1884, even though the species collected on three earlier occasions by other people. In 1888, Medley Wood, the curator of the Durban Botanical Garden sent plants to Kew, which subsequently flowered. This is native to South Africa, Madagascar, Asia, South America and Indonesia.

The first official description of South African species G. jamesonii was made by J.D. Hooker in Curtis Botanical magazine with coloured illustration in 1889. An active society on this subject is to be found in Australia under the name W.A. Gerbera Society. The society has used a classification system of gerbera. The breeding of gerbera started at the end of the nineteenth century in Cambridge, England, when two South African sp. were crossed. G. jamesonii native to Natal and Transvaal and G. viridifolia from western Cape were crossed together by R. Irwin Lynch in 1887, after both these species were introduced to Cambridge Botanic Gardens by Jameson and the hybrid was named as G. x cantabrigensis.

Abstract: Gerberas are popular seed and vegetatively propagated flower crop with numerous phenotypes, series and cultivars available on the world market. Crop improvement is currently focused on bicoloured mini gerbera, semi-double and spider type forms, and hybrid cultivars with expanded colour range to boost productivity and profitability. Application of genetic engineering techniques and tools are aiding in modern gerbera breeding. The need for new cultivars remains strong as production system become more specialized and new markets develop. Breeding objectives with high priority include cut flower cultivars highly resistant to diseases, tolerant to low temperature, and floral longevity.

INTRODUCTION

Gerbera is a herbaceous flower crop with leafless stalk and daisy like flowers. It is popularly known as Barberton daisy, Transvaal daisy, Veldt daisy, African daisy and Hilton daisy. Gerbera is a popular cut flower grown throughout the world. It is ideally suited for flower beds, borders, pots, window boxes and gardens. The gerbera plant is also used in the preparation of traditional Chinese medicine i.e. Tu-er-feng, derived from whole plant of gerbera, is used for curing cold with cough and for rheumatism. Gerberas have wide range of bright, vivid and clear colours. Some of the flower colours available today are pink, yellow, red, scarlet, orange, salmon, lavender, violet, rose, cream and white. They are great plants for adding colour to any room or garden to create festive and elegant display. These spectacular, eye catching flowers are a complement to any kind of flower arrangement or bouquet and have a longer vase life. Their daisy shaped flowers have a velvet texture with green or dark centers and are cherished by florists and the cut flower industry.

It grows well in open tropical and sub tropical regions but in a temperate climate should be protected from frost and cultivated in greenhouses. Production of quality flowers of gerberas can be done under protected conditions, since they require partial shade. In California and Florida, it is grown in outdoors and is becoming increasingly popular as potted plant and bedding plant, whereas in other parts of United States it is cultivated in greenhouses. Gerbera cultivation has emerged as a very important option to progressive farmers in many parts of India, especially in Maharashtra, Karnataka, north-eastern states, U.P. and Uttaranchal.

This chapter introduces Gerbera as a popular cut flower known for its daisy-like blooms, wide range of colours and long vase life. It examines its origin, distribution, genetics, botanical characteristics and commercially important varieties. The chapter discusses greenhouse production, environmental requirements, propagation, irrigation, nutrition, cultural practices, pest and disease management, harvesting and postharvest handling. It emphasizes the importance of protected cultivation for producing high-quality gerbera flowers in different agro-climatic regions of India

Gerbera, commonly known as Transvaal Daisy, Barberton Daisy or African Daisy, is cultivated throughout the world under a wide range of climatic conditions for its attractive flowers. It is highly suitable for beds, borders, pots and rock gardens. The wide range of colour and the attractive shape of flowers suit very well in flower arrangements. The cut blooms have long vase life. Today Gerbera is known as an important commercial flower crop and belongs to the most important plant species in the world, together with the rose, chrysanthemum, carnation and tulip. Gerbera is amongst the ten most important commercially grown flower crops in the world.

Introduction

Gerbera is a herbaceous flower crop with leafless stalk and daisy like flowers. It is also known as Transvaal daisy. Gerbera is a popular cut flower grown throughout the world. It is ideally suited for flower beds, borders, pots and gardens.

In California and Florida, it is grown in outdoors and is becoming increasingly popular as potted plant and bedding plant, whereas in other parts of United States it is cultivated in greenhouses. Gerbera cultivation has emerged as a very important option to progressive farmers in many parts of India, especially in Maharashtra, Karnataka, North Eastern states, U.P. and Uttarakhand.

Genetics

The genus Gerbera belongs to family Asteraceae. Gerbera comprises about 70 species, which are of African and Asiatic origin. The basic chromosome number is n=25. G. jamesonii is a diploid species with somatic chromosome number 2n=50. Among the different species, G. jamesonii is the only species under cultivation.

Wild relatives of cultivated plants constitute a part of crop genepool, which possess genes that have great potential for their utilization in crop improvement programmes. Wild genepools especially those occurring in biotically disturbed habitats are under threat of genetic erosion and require immediate collection to make use of their wider adaptability, tolerance/resistance to disease, insect-pests, yield, quality attributes and other biotic and abiotic traits. The wild relatives of crop plants occur in all the major phytogeographical regions of India as component of different vegetation types and exhibit variable life forms and habitat specificity. High priority demand is towards collection of those that are at the risk of erosion due to habitat disturbance by manmade/ natural causes and over-exploitation of species for commercial use.

The centres of endemism are also centres of genetic diversity and consequently centres of speciation. Unless suitable conservation methods are applied, important wild relatives of crop plants and rare endemic species in particular have poor chances of survival. Based on the efforts made in the past for collection of diversity in wild relatives of crop plants priorities have been recommended for their conservation on the basis of species status (endemic, rare/threatened/endangered, etc.) and those that have high demand in crop improvement programmes (Arora and Nayar, 1984; Nayar, 1996, 1997; Wiersema and Leon, 1999; Hanelt and Institute of Plant Genetics and Crop Plant Research, 2001).

All economically important species at the risk of genetic erosion require attention for collection and conservation. Highest priority may be assigned to those having highly specialized habitat requirement and those that are endemic/rare/endangered to Indian region. Species producing seeds in large quantity and having easy method of multiplication or conservation are at lower risk than those producing less seed and of the recalcitrant or orthodox types where the storage protocols have been least worked out. Species having short seed viability or low germinability need special investigation.

Wild relatives of cultivated plants constitute a part of crop genepool, which possess genes that have great potential for their utilization in crop improvement programmes. Wild genepools especially those occurring in biotically disturbed habitats are under threat of genetic erosion and require immediate collection to make use of their wider adaptability, tolerance/resistance to disease, insect-pests, yield, quality attributes and other biotic and abiotic traits. The wild relatives of crop plants occur in all the major phytogeographical regions of India as component of different vegetation types and exhibit variable life forms and habitat specificity. High priority demand is towards collection of those that are at the risk of erosion due to habitat disturbance by manmade/natural causes and over-exploitation of species for commercial use.

Plant genetic resources are the key components of crop improvement because of development of precision breeding system through biotechnological approaches as well as conventional system of breeding like selection and hybridization. Breeding system of hybridization is replaced by cybrid technology that facilitates transferring more qualitative and quantitative traits rapidly within incompatible genotypes. Aonla is mainly recognized for its high nutritive, medicinal and therapeutic properties. It contains high amount of vitamin-C (500-1500mg/100g), it is also rich in pectin, iron, calcium and phosphorus. Dried fruits of aonla can be used in curing haemorrhage, diarrhea, chronic dysentery, diabetes, jaundice, dyspepsia and cough. Recently, aonla gained more popularity in arid and semi- arid region of the country because of its hardy nature, prolific bearing capacity and ability to grow under various adversities as a remunerative crop (Shukla et al.2002). Aonla is the main ingredient of Chavanprash and Triphala powder, used for curing different   human ailments. Fruit contains a chemical substance gallic acid and leucoanthocyanin, having antioxidant property.

As mentioned above, all the present day Bougainvillea cultivars being grown in India are either exotics introduced from abroad (Africa, USA, U.K., South Africa, Australia, Philippines, Sri Lanka, Jamaica, Canary Island, Singapore, Java, West Indis etc) or developed by concentrated efforts of nurserymen, individuals or scientific institutions. Bougainvillea are widely grown in Indian gardens. There are numerous varieties and cultivars that are important from floriculture point of view. Most of the present day varieties have arisen through bud sports, chance crossing and induced mutation. India is one of the major repository of wide range of genotypes of Bougainvillea and is perhaps the foremost among nations which have done appreciable good work on Bougainvillea. The genetic variation in plants is considered to be an unlimited resource. The main aim of germplasm collections are assemblies of genotypes or populations representative of cultivars, genetic stocks, wild species etc. which are maintained in forms such as plants, seeds and tissue cultures. The primitive cultivars and wild relatives of crop plants constitute a pool of genetic diversity.  The  germplasm collection and conservation of plant genetic resources provide the diverse raw materials for immediate or future breeding programme. Activities that relate to the conservation and use of plant genetic resources include : exploration and collection; characterization and evaluation; conservation; assessment of variation and identification of useful genes; and exchange and genetic enhancement. Germplasm are the basic raw materials, which after proper screening can be utilized either for direct commercial exploitation or can be used in breeding programme for developing new and novel varieties.

Giant granadilla, Passiflora quadrangularis L. (syn. P. macrocarpa M.T. Mast.) is the largest fruit in the genus Passiflora. With more than 450 species, Passiflora is the most important genus of family Passifloraceae. It is further divided into 24 subgenera (Killip, 1938). Most of the species producing an edible fruit belong to subgenera Passiflora and Tacsonia. Many species have ornamental interest because of unique and spectacular flowers, or are used for their sedative, antispasmodic, antibacterial, and insect-resistant properties.

Gladiolus (Gladiolus spp.) belongs to family Iridaceae and subfamily Ixioideae. Gladiolus is a Latin word, the diminutive of “gladius” meaning sword, sometimes called as sword lily. The plant is herbaceous perennial and semi-hardy. It is commonly grown from rounded, symmetrical corm, which is enveloped in several layers of brownish, fibrous tunics. The cultivated gladiolus is a complex assemblage of cultivars originating from complex ancestry of natural selection or hybridization involving several species. Modern garden gladiolus is a complex of at least twelve species (Raghava, 2000). It is an important commercial cut flower in both domestic and international markets.

ORIGIN AND DISTRIBUTION

There are two main centres of distribution of Gladiolus species, one in Mediterranean basin and the other in southern and central Africa (Lewis et al., 1972 and Delpierre and Du Plessis 1974). Most of the species are native to Mediterranean region and tropical parts of the South Africa, particularly the region of the Cape of Good Hope (Raghava, 2000).

The name gladiolus was coined by Pliny the Elder (A.D. 23-79) to describe the shape of the leaf which resembles that of sword (Latin word gladius meaning sword). It is native to South Africa. Most of the species scattered throughout South Africa, Zambesi River, Rhodesia, Mt. Kilimangero, Tanganyika, Natal and Cape of Good Hope, Republic of South Africa, Madagascar and Abyssinia (Ethiopia). Gladioli are said to be cultivated since the days of ancient Greece. As evidenced by a record in Lyte’s Nieeve Herball, it is known since 1578, first introduced in France and soon after spread to England, Germany, Holland and North America. No evidence is available that any species of gladiolus of South African origin was ever grown in Europe prior to 1700. Cornutus described Gladiolus aethiopion bearing light red-scarlet flowers and is from Cape region. In the middle of eighteenth century the South African species were introduced into Europe. Gladiolus tristis closely related to G. grandis was first introduced in Chelsea Physii Garden, UK in 1745 from Cape region and Walat. G. illyricus is native to Mediterranean region. In 1749, Gladiolus grandis was naturalized in Spain from South Africa.

By 1816, France, Holland and Belgium started raising hybrids, the first being Ghant Gladiolus, Gladiolus gandavensis, in 1841, by crossing G. cardinalis and G. psittacinus and vice-versa. Gladiolus primulinus having central petals with hooded characteristics was collected from its native place on the banks of Zambesi River near Victoria Falls in 1902. Victor Lemoine of France used G. primulinus for producing ‘Primulinus Hybrids’ during 1906-1908 while A.E. Kunderd of Indiana, U.S.A. in 1915 produced a similar race ‘Primulinus Grandiflora Hybrids’ by using same parentage as Lemoine. In 1907, first ruffled gladiolus with fimbriated and frilled petals was produced by the efforts of Kunderd.

Gladiolus is a potential flower crop for floriculture industry in India. Gladiolus is commercially grown in West Bengal, Maharashtra, Uttar Pradesh, Uttarakhand, Punjab, Haryana, Sikkim, Jammu and Kashmir, Karnataka, Gujarat, Himachal Pradesh, Tamil Nadu, Madhya Pradesh, Delhi and Rajasthan. The area of the states of U.P., Uttarakhand, Haryana, Punjab, Rajasthan and H.P. and surrounding Delhi grow gladiolus mostly for Delhi flower market (Singh, 2006).

Abstract: Gladiolus, commonly known as ‘sword lily’ is native to South Africa. Most members of genus are heteroploids. It has pivotal place as cut flower with novel phenotypes (Grandifloras, Primulinus, Butterfly, Miniature, Face Ups, Orchideola), cultivars available on world market. Modern types are evolved through conventional breeding and very few through mutation breeding. Their continued popularity has warranted extensive breeding and development of fragrant, winter hardy and disease resistant cultivars. Breeders have reported close linkages in variety of traits. Cross pollinating nature coupled with high heterozygosity makes heredity transmission unpredictable. Future crop idiotypes are proposed to continue transformation of this important crop during the next millennium.

Gladiolus is a major commercial cut flower appreciated for its long spikes, attractive florets and wide range of colours. This chapter describes its origin, species, genetics, botanical characteristics and improved varieties. It explains corm and cormel propagation, climate, soil, planting time, spacing, nutrient management, irrigation, staking and other crop-management practices. The chapter also discusses major pests and diseases, physiological problems, harvesting stages, grading, postharvest handling, storage and yield. Emphasis is placed on producing long, straight and high-quality spikes suitable for domestic and export markets. The chapter provides practical production guidance for commercial gladiolus cultivation under Indian conditions. effectively.

Gladiolus, commonly known as ‘Sword lily’ which derives its name from latin word Gladius meaning sword shape foliage. The name gladiolus was originally given by Pliny the Elder (A.D.23-79). It is an important florist crop, cut flower in the domestic and international flower trade. It is relatively easy to grow, suitable for cut flower, pots, hanging basket, bedding and exhibition. An engrossing cut spikes bear a large number of florets on rachis, which exhibit various sizes, forms, with smooth, ruffled, deeply crinkled or laciniated tepals. As a genus it was first proposed by Tournefort and which was accepted in botanical litreture by Linnaeus through his description “Species Plantarum” in 1753 and “Genera Planterum” in 1754. The gladioli are said to be cultivated since the days of ancient Greece. The genus Gladiolus, at present comprises with 255 species (Mishra et al.,1995; Goldblatt and Manning,1998) with wide distribution occurs in Central Europe, Mediterranean region, Central and South Africa are primary centre of origin. Campbell and Bower (2003) mentions about one more species Gladiolus scabridus- a new species 1978 from the mountains of northern Kwa Zulu- Natal and southern Swaziland, restricted to well- drained rocky habitat and where it is endemic. This species has enormous commercial potential have large number of pink flowers. So now number of recorded gladiolus species are more than 260. The gladiolus was brought into cultivation first during Greek period. Gladiolus has been documented in literature since 1578, as evidenced from the record Lyte’s Nievve herbal. This was first time introduced in France and soon after spread to England, Germany, Holland and North America. In India, its cultivation dates back to 19^th century as mentioned in the ‘Firminger’s Manual of Gardening in India (1863)’ that Mr. Charles Gray of Conoor grew some gladioli from corms and seeds at his garden. The European Union gladiolus importing countries are Germany, Belgium, Luxembourg, Denmark, Spain, France, United Kingdom, The Netherlands, Italy and Portugal.

INTRODUCTION

Gladiolus is very popular bulbous flowering plant. It is an important commercial flower crop and having pivotal place as cut flower both in domestic and international market. Gladiolus was introduced into cultivation towards the end of the sixteenth century. It is relatively easy to grow and is ideal for bedding and exhibition. The fascinating spikes bear a large number of florets, which exhibit varying sizes and forms with smooth, ruffled or deeply crinkled tepals. Except true blue and green, practically all colours are available in gladiolus. The flowers are used in flower arrangement, in bouquets and for indoor decorations. Now it ranks next to tulip in the Netherlands and other European countries in trade for use as cut flowers of bulbous crops and fourth in international trade of ornamental for cut flowers. Its keeping quality makes it a very popular commercial cut flower after rose. Gladiolus occupies first place in terms of returns as compared to other cut flowers. Gladiolus, a potential floral crop forms part of Rs. 200 crores floriculture industry in the country. It is estimated that throughout India gladiolus is grown in about 1270 ha, producing about 127 million cut spikes. In India, gladiolus is commercially grown in West Bengal, Maharashtra, Uttar Pradesh, Uttaranchal, Punjab, Haryana, Sikkim, Jammu and Kashmir, Karnataka, Gujarat, Himachal Pradesh, Tamil Nadu, Madhya Pradesh, Delhi and Rajasthan. The area of the states of U.P., Uttaranchal, Haryana, Punjab, Rajasthan and H.P. and surrounding Delhi grow gladiolus mostly for Delhi flower market. Important pockets of commercial gladiolus cultivation are Lucknow, Meerut, Bulandsahar, Ghaziabad and Uttaranchal. In Delhi market, gladiolus is available virtually throughout the year as growers stagger the corm planting from July to December to harvest the flowers from September to May. During the period between June to December, the supply of gladiolus spikes to Delhi market comes from Jammu and Kashmir, Uttaranchal and Himachal Pradesh. The high remuneration in this market is obtained from winter grown gladiolus.

Gladiolus is a flower of breathtaking beauty with a wide range of colours, size and form. Its magnificent inflorescence has won for it, a place of pride in gardens and value as a cut flower. Except true blue and green practically all colours are available in gladiolus. Flowers are long lasting and stand transport well. They are widely used in flower arrangements, bouquets and indoor decorations. It is estimated that gladiolus is grown throughout India over on area of 1270 ha (Arora et al. 2002).

Introduction

Gladiolus is an important commercial flower crop and is very popular as cut flower both in domestic and international market. It is a bulbous plant. Gladiolus was introduced into cultivation towards the end of the sixteenth century. The flowers are used in flower arrangement, in bouquets and for indoor decorations. In India gladiolus is commercially grown in West Bengal, Maharashtra, Uttar Pradesh, Uttarakhand, Punjab, Haryana, Sikkim, Jammu and Kashmir, Karnataka, Gujarat, Himachal Pradesh, Tamil Nadu, Madhya Pradesh, Delhi and Rajasthan.

Genetics

Gladiolus commonly called Sword Lily belongs to family Iridaceae and subfamily Ixioideae. There are about 260 species of gladiolus. It has the basic chromosome number n=15. Most members of genus are heteroploids having very small chromosomes ranging from 2n=30 to 120 (Diploid, triploid, tetraploid, pentaploid, hexaploid, octaploid and hyperaneauploid).