eChapters

Introduction

The horticulture sector has been a driving force in stimulating a healthy growth trend in Indian agriculture. This sector contributes around 28 per cent of the GDP from about 13.08 per cent of the area and 37 percent of the total exports of the agricultural commodities (Rather et al., 2013)  India is currently producing 257.2 million tonnes of horticulture produce from an area of 23 million ha. Over the last decade, the area under horticulture grew by about 3.8% per annum but production rose by 7.6% per annum. The higher growth rate in horticulture was brought about by improvement in productivity of horticulture crops, which increased by about 28% between 2001-02 and 2011-12. (Anonymous, 2012)

The main purpose of herbal drug technology is to convert botanical materials into medicines with a focus on standardization and quality control with proper integration of modern scientific techniques and traditional knowledge. The use of chromatographic techniques and marker compounds to standardize botanical preparations has limitations because of their variable sources and chemical complexity. DNA-based molecular markers have utility in fields like taxonomy, physiology, embryology, genetics etc. DNA based techniques have widely been used for authentication of plant species of medicinal importance. Pharmacognosy mainly addresses the quality related issues using routine botanical and organoleptic parameters of crude drugs, and chemoprofiling assisted characterization with chromatographic and spectroscopic techniques. The new pharmacognosy includes all the aspects of drug development and discovery where the biotechnology driven applications play an important role. Current focus on chemotype-driven fingerprinting and related techniques require integration with genotype-driven molecular techniques so that an optimal characterization of botanical materials is possible. This review provides a brief account of various DNA based technologies that are useful in genotyping and quick identification of botanicals with suitable examples.

Abstract

Tea [Camellia sinensis (L.) O. Kuntze], of family Theaceae, is an important cash crop  worldwide. It is an evergreen shrub cultivated in humid and sub-humid tropical, sub-tropical, and temperate regions of the world, and grows mainly on acidic soils. Due to limitation of expansion of tea cultivation in new land, import of low cost high quality tea, increasing vertical productivity is the best option of Indian tea Industry. The primary need of the tea industry today is to develop high yielding clones with better quality and stress tolerance potential. Conventional tea research has contributed appreciably to the genetic improvement of tea over the past several decades, but it is time-consuming and labor intensive.

It is difficult to define ‘stress’ precisely. In engineering and the physical sciences, it is usually defined as the force applied per unit area. But as far as in biology, stress it is usually described more loosely as any factor that disturbs normal functioning of an organism. Any environmental factor potentially unfavorable to plant is termed as stress.

The effect of stress on plant condition is called strain. Stress is the action, whereas strain is the reaction. Stress may be defined as external conditions that adversely affect growth, development and productivity. Stress may further classified as:

• Zero stress: This is the level of exposure of plants to environmental factors that leads to neither injury nor reduction in growth and yield of crops.
• Stress escape: Capacity of the plant to complete its life cycle during wet season before onset of stress.

English name : Bishop’s weed, Ajowan
Botanical name : Trachyspermumammi (L.) Sprague
Family : Apiaceae
Chromosome number : 2n = 18
Centre of origin : Egypt
Ajowan is an annual herbaceous plant grown for its herb and greyish brown fruits (seeds) which constitute the spice. The trade name ajowan or ajwain
is based on the Indian name which is derived for adarjawan. It is known as ‘Bishop’s weed’ in English. Ajowan is grown for its seeds (fruits) the composition of which is as follows; moisture 8.9%, protein 15.4%, fat (ether extract) 18.1%, crude fibre 11.9%, carbohydrates 38.6%, mineral matter (total ash) 7.1%, calcium 1.42%, phosphorus 0.30%, iron 14.6 mg/100 g, vitamins (per 100 g) thiamine 0.21 mg, riboflavin 0.28 mg, nicotinic acid 2.1 mg, carotene 71 mg. Its calorific value is 379.

Anthracnose
Diagnostic Symptoms
• Pathogen attacks all aerial parts of plants and at any stage of their growth.
• Symptoms are circular, black, sunken spots with dark center and bright red orange margins on leaves and pods.
• In severe infections, the affected parts wither off.
• Seedlings get blighted due to infection soon after seed germination.
• Infected pods have discolored seeds (Fig.3).
Etiology
Scientific Classification
Kingdom : Fungi
Division : Eumycota
Sub- division : Deuteromycotina

19.1. Origin and distribution

• Native to Eurasia and introduced in America, Australia and South Africa.

• Grown throughout dry parts of India

19.2. Economic importance

• Whole plant - Useful for rheumatalgia, swellings, cough, asthma, bronchitis, wounds, ulcers, flatulence, dyspepsia, hepatomegaly, otalgia, hiccough, opthalmopathy, vomiting

• Used in cardiopathy, leprosy, skin diseases, fever, splenomegaly, emollient

• As an antiseptic, anti-inflammatory, expectorant, anodyne, vulnerary, digestive, laxative, diuretic, cardiotonic, depurative, diaphoretic, febrifuge, rejuvenating, sedative, alterant and tonic.

19.3. Climate

• Well in tropical and subtropical climate

1.0 INTRODUCTION

Black pepper popularly known as ‘King of spices’ or ‘Black gold’ is one of the important and earliest known spices produced and exported from India. Its original home is the dense evergreen forests of Western Ghats in south India. The flavour and fragrance of this oriental spice had a magic spell in human civilization and culture since very ancient days. Expeditions in search of pepper led to the discovery of new trade routes.

English name : Black pepper
Botanical name : Piper nigrum L.
Family : Piperaceae
Chromosome number : 2n = 52
Centre of origin : Western Ghats, India
Black of the commerce is the dried mature fruit of the perennial climbing vine of Piper nigrum L. It is one of the oldest and most significant spices in the world. Black pepper refers to the whole dried fruit, which is still unripe, while white pepper is made from the fruit after the mesocarp has been removed. Among all spice crops, pepper holds the highest share in foreign exchange, earning it the title of the “King of Spices.” An analysis of various types of black pepper from Kerala, Karnataka, Tamil Nadu and Assam revealed the following ranges of values: moisture content (8.7-14.1%), total nitrogen (1.55-2.60%), nitrogen in non-volatile ether extract (2.70-4.22%), volatile ether extract (3.0-4.2%), non-volatile ether extract (3.9-11.5%), alcohol extract (4.4-12.0%), starch (28-49%), crude fibre (8.7- 18.0%), crude piperine (2.8-9%), piperine (spectrophotometrically) (1.7- 7.4%), total ash (3.9-5.7%), and acid-insoluble ash (0.03-0.55%). Starch is the principal constituent of pepper, making up 34.8% of black pepper, 56.5% of white pepper, and 63.2% of decorticated white pepper. Pepper starch consists of small, polygonal granules similar to rice but much smaller. The alkaloid piperine (melting point: 29-30°C) is the primary compound responsible for black pepper’s characteristic biting taste. Other pungent alkaloids present in smaller quantities include chavicine, piperidine, and piperetine. 

Black pepper (Piper nigrum L.), famous as “Black Gold” and also known the “King of Spices” is the most important agricultural commodities of commerce and trade in India. It is the most important, most popular and most widely used spice in the World. Of the total spices traded internationally, black pepper accounts for 34 per cent. South West India is the traditional home of this important spice, particularly the Malabar Coast. Black pepper is also grown in tropical zones such as the Asia Pacific Region, mainly India, Indonesia, Malaysia, Sri Lanka, Thailand, China, Vietnam and Cambodia. Outside the Asia Pacific region, the crop is distributed in Brazil, Mexico, Guatemala, etc., totaling about 26 countries. It is grown successfully between 20°N and 20°S latitude and from mean sea level to 1500 meters above sea level. World’s black pepper production varies in the range of 2.7 to 3.35 tonnes annually. The major production share is contributed by Vietnam with an annual production of 80,000 tonnes. About 85 per cent of the World trade of pepper is determined by the largest pepper producing countries in the International Pepper Community (IPC). World export fluctuates around 1.5–2.5 lakhs tonnes per annum. Global black pepper supply has shown a drastic increase over last ten years except in 2008–09. Vietnam’s sudden increase in production has resulted in the massive global out put of 3–3.5 lakhs metric tonnes from 1.9–2.0 lakhs metric tonnes in late nineties. 

Some species of turmeric have been documented from China, Australia, and the South Pacific, despite the fact that turmeric has been farmed all throughout the tropics. Turmeric along with the 110 species is found growing in Tropical Asia, with India, Myanmar, and Thailand having the most diversity. Black turmeric (Curcuma caesia), along with Curcuma aromatica, Curcuma amada, Curcuma aeruginosa, Curcuma longa, and Curcuma zanthorrizha, is one of the economically important species native to India. Curcuma caesia (black turmeric/ black zedoary) is a critically endangered perennial herb in the Zingiberaceae family native to India. Traditional healers employ the Curcuma genus to treat a variety of diseases in ethno medical traditions, but Curcuma caesia Roxb. is a relatively unknown and unexplored medication. In the Indian subcontinent, black turmeric is frequently used as a spice, food preservative, and colouring ingredient. Black turmeric is the most recent spice to be chemically studied, with a few papers describing 100 or more secondary metabolites from this plant, and it will continue to be a source of new bioactive chemicals.

Distribution

Black turmeric is a native of Northeastern and Central India, where it has long been used in traditional rituals and medicinal treatments. Within an altitudinal range of 200–1000 m, the species is found in moist deciduous woods, primarily in Bengal, North East, and Central India. It develops as a woodland ground cover in subtropical to temperate climates.

Abstract

Blumea lacera (Burm f.) DC. belonging to family Asteraceae is an annual herb with a strong odour. This herb is distributed throughout the plains of North-west India upto an elevation of 2,900 m and is also reported from Tropical Africa, South East Asia, Bangladesh, Nepal, Indonesia, Malaysia, Philippines, Thailand and Vietnam. Blumea lacera is a plant with camphor-like odour, and is highly variable, grows in grassland, field, roadside and forest edge. Traditionally, it is used internally and externally as a hemostatic and anti-inflammatory agent. The juice of leaves along with black pepper (Piper nigrum) is given to cure bleeding piles. This herb is also offered as anthelmintic (especially for threadworm). Leaves and roots are used as astringent, diuretic and febrifuge and even in cuts and wounds. Bruises and ulcers can also be treated by applying fresh juice or extract. For treating piles (Hemorrhoids), the leaves of B. lacera has been used in ethnomedicinal practices.The phytoconstituents present in leaves are mainly terpenoids, glycosides, volatile oils, some glycoalkaloids and many more yet to be determined. B. lacera has been used as an ingredient in many pharmaceutical preparations. The chapter gives a brief idea about the ethnomedicinal usage, phytochemical profile, pharmacological usage and the future perspective of B. lacera. and would help the researchers for better understanding of this plant as well as developing plant derived molecules in drug discovery process.

ABSTRACT

Boswellia serrata Roxb. ex Colebr. known as ’Indian Frankincense’ is a multiferous medicinal plants known for boswellic acid and it has application in traditional ayurvedic formulation. The plant possess anti-rheumatic, anti-pyretic, anti-cancerous, anti-inflammatory, anti-hyperlipidemic, anti-coronary, analgesic and hepatoprotective properties due to the presence of several loaded bioactive compounds such as α-amyrins, β-boswellic acid, acetyl-β-boswellic acid, 11-ketoβ-boswellic acid, acetyl-11-keto-β-boswellic acid, tetracyclic triterpenoic acids and several others in minor quantities in different parts of plants. Mostly oleo-gum resin is usually used in Indian system of medicine, and acetyl-11-keto-β-boswellic acid (AKBA) is the most potent 5-lipoxygenase enzyme inhibitor responsible for inflammation, anti-cancer (G1 phase arrest) and other group of enzymes liable for allied activities. Besides, it yields essential oils and its major constituents are monoterpenoids such as pinene (2.05-64.7%), cisverbenol (1.97%), trans-pinocarveol (1.80%), borneol (1.78%), myrcene (1.71%), verbenone (1.71%), limonene (1.42%), thuja-2,4(10)-diene (1.18%) and p-cymene (1.0%). The copaene is the only sesquiterpene reported in minor quantities from essential oils of this species. Its essential show different actvities in relation to anti-cancer and antiinflammatory. From future perspectives, there is a need to carry out more research on its constituents for the development of value added products and drugs. Investigation on ecology and application of biotechnological intervention would helps in varities improvememnt, cultivation and conservation of this species.

In previous Chapter, a list was furnished mentioning common names, botanical names, family and types of the all plants mentioned in the Ramayana. The purpose was to know the total number of plants and trees mentioned besides providing reference of the Shlokas as well. This information led to identification of the plants and trees but details about their morphological and floral characters were not mentioned. Therefore, in this Chapter details of the trees and other plants have been provided. This will help to know all about their botanical standing, conservation status, nativity, morphological and floral characters so that one can know the full details including present-day status, availability and importance (Cowen, 1950; Randhawa, 1983; Roy, 2015; 2024).

1. Botanical Name: Adenanthera pavonina L.

Synonym: Adenanthera gersenii Scheff.

Common Name: ‘Red Bead Tree’, ‘Curly Bean’, ‘Red Sandal Wood’

Family: Fabaceae

Nativity: India, Sri Lanka, Southeast Asia to Queensland

Conservation Status: Common (Not Threatened)

Type: Tree

Morphological and Floral Characters: Red Bead Tree is a timber tree found wild in India. Leaves compound, bipinnate, green when young, turning yellow on maturity. Flowers small, yellowish, grows in a dense, drooping almost cat tail like flower-heads. Fruits curved pods, hanging, green that turn brown, coil up and split open on ripening, showing small bright red seeds which are attractive, used as beads in jewellery.

Pyrethrum is the most widely used botanical insecticide. The active ingredient, pyrethrin, is extracted from a chrysanthemum plant, grown primarily in Kenya, Rwanda, Tanzania and Ecuador. Pyrethrum is the powdered, dried flower head of the pyrethrum daisy, Chrysanthemum cinerariaefolium.

Botany

Several vernacular names of kewda are reported in different languages all over the world ; viz.

Keora, Ketgi, Keura, Gagandhul in Hindi; Keya, Ketaki, Kea, Ketkikea in Bengali; Kiya, Kia in Odia; Kewoda in Gujrati; Ato, Keto, Covasso in Konkani; Keoda, Keora in Marathi; Talemara in Kannada; Gedaji, Gedangimogali, Gojjangi, Ketaki, Mugali, Mogali in Telugu; Kaidai, Talhai, Thasai, Thazh, Kechiya, Kedagi, Kedagai, Muchali in Tamil; Kaita, Ketaki, Palliyambu in Malayalam; Gandhapuspa, Dalapushpa, Shivadvishta, Ketaki, Suchika pushpa in Sanskrit; Leram in Nicoberese; Mudukeyiya, wetakey in Sinhalese; Keora in Urdu; Gulkiri, Kader, Kadi in Persian; Satthapu, Tsatthapu in Burmese; Nalu in Nepalese; Togenashiadan in Japanese; Kadar, Kadhi, Kazi in Arabic Kaide, Kedige, Ketaki, Gedege, Mundige in Canarese; Kedgi in Deccan; Vaquois odorant in French and Schrauben palme in German. (Kirtikar and Basu, 1935; Randhawa and Mukhopadhyay, 1986; Kapoor, 1990).

18.1. Origin and distribution

• Native of wetlands of Southern and Eastern India, Australia, Europe, Africa, Asia, and North and South America

• Cultivated throughout tropical and subtropical India.

18.2. Economic importance

• Economic part - Whole plant

• Chemical constituents: Bacosides

• Used as a nerve tonic and for epilepsy and insanity.

• Used as diuretic and for treating rheumatisim, asthma and hoarseness.

Horticulture includes variety and diversity of crops. Thus, it can be divided into several branches depending upon the crops it deals with. Following are the branches of horticulture.

1. Pomology: The term pomology is a combination of two Latin words pome i.e. fruit and logos i.e. science. Recently it has been designated as ‘Fruit Science’. It deals with the study of fruit crops. It includes all aspects of fruit crops e.g. importance, uses, origin and distribution, area and production, soil and climate requirement, botany, nursery raising, cultivation practices, plant protection measures, stage and methods of harvesting, post-harvest handling, marketing, export, import, etc. e.g. mango, banana, citrus, grape, apple, peach, pomegranate, fig, etc.
2. Olericulture: The term olericulture is also derived from the Latin word. Cultivation of vegetable crops. Recently it has been designated as ‘Vegetable Science’. Vegetables are said to be herbaceous plants of which edible plant parts are used for consumption either cooked or salad. It may be stem, root, leaf, flower, etc. It also includes all aspects of vegetable crops e.g. importance, uses, origin and distribution, area and production, soil and climate requirement, botany, nursery raising, cultivation practices, plant protection measures, stage and method of harvesting, post-harvest handling, marketing, export, import, etc. e.g. tomato, brinjal, chilli, cucumber, red pumpkin, watermelon, cabbage, cauliflower, etc.
3. Ornamental Horticulture: It includes various groups of ornamental plants used to decorate in-door and out-door areas. Ornamental horticulture is the study of growing, arranging and tending decorative plants and flowers. It is further divided into.

Morphological characteristics

Caesalpinia sappan is a thorny, small to medium-sized plant with prominent spines on the stem and leaf rachis that grows up to 10 metres tall. Branchlets are lenticellate, dull, and usually have paired, recurved stipular prickles. With 9–14 pairs of pinnae, the leaves are big, hairy, or glabrate.

Floral characteristics

The inflorescence is a supra-axillary and terminal raceme with golden yellow flowers that creates a huge panicle. The fruit is an elliptical pod that ranges in colour from brown to black. The peak flowering and fruiting season is from July to September, with November being a rare exception.

Distribution

The crop is native to the Indo-Malayan region. The species can be found throughout the Indian peninsula. It is also produced as a plantation crop in all of the Southern states, as well as in Odisha and Bengal to the north.

Breadfruit (Artocarpus altilis) belongs to family Moraceae. The Latin name is derived from Greek words (artos = bread, karpos = fruit), and altilis meaning ‘fat’. Baked or roasted in a fire, the fruit has a starchy texture and fragrance, reminiscent of fresh baked food.

Breadfruit is an important staple crop and component of traditional agro-forestry systems in the Pacific for more than 3,000 years. This species originated in the South Pacific and spread throughout Oceania by intrepid islanders settling the numerous islands of Melanesia, Micronesia and Polynesia. Due to efforts of Captain Bligh and French voyagers, a few seedless varieties from Polynesia were introduced to the Caribbean in the late 1700s. These gradually spread to other tropical regions. Breadfruit is now grown in close to 90 countries.

No breeding work has so far been done on kewda except selection of some promising morphotypes. Varietal improvement in kewda by breeding poses problems mainly due to dioecious nature and long life cycle of the plants, besides heterozygosity. Lack of promising strains is a major drawback. The wild growing population of kewda has a measure of variation and this has never been documented and utilized, which naturally offer a good scope for improvement of kewda by resorting to selection and subsequently propagating them vegetatively. Since a wide range of variation is available and the plants are mostly propagated by vegetative means, it is worthwhile to collect genetic stocks, access variability in morphological characters and attempt a breeding programme after selecting the desirable types for high flower production as the ultimate economate economic value of the plantations depends on the quantity of flower production to be used industrially in the distillation units.

The current status of advances in fruit breeding is through the development of different scientific approaches augmented by ever-increasing knowledge and application of genetics, statistics and biotechnology. These innovative approaches very specific for a species in most of the cases, are the major tools of the fruit breeders to tackle the different objectives of fruit improvement programme at present Fruit improvement is a very slow process due to the long generation cycle varying from species to species, thereby reducing the chances of exploiting genetic recombination. Besides this factor, most of the fruit species are either highly heterozygous or polyploidy in nature. Due to this reason the asexual propagation of fruit plants is a conventional means while wide segregation occurs upon sexual reproduction. Such expectation is not unlikely because the types selected as commercial varieties are definitely to be vigorous ones, and a positive correlation exists between vigour and heterozygosity.

This discussion on breeding of fruit plants involves different aspect of the species concerned is so essential as well as fundamental that discussion of them must procede any consideration of breeding methods themselves. The plant species are grouped into two-self-pollinated and cross-pollinated. These two groups have emerged on the basis of pollination mechanism which plays significant role in the method of breeding application. To emphasize the difference in the reproductive systems of these two groups one should know the details about floral biology, mechanism of pollination control, reproduction by asexual and sexual means and self and cross-pollinated plants.

29.1 Introduction

An environmental factor that interferes with the complete expression of genotypic potential of a crop is called stress. The stresses may be biotic (pathogens, insect-pests, etc.) or abiotic. The main abiotic stresses are drought, mineral deficiency/toxicity and heat or cold. The losses due to abiotic stresses can be minimized by:

Crop management: The degree of stress can be reduced by suitable agronomic/soil management practices, e.g., irrigation to relieve drought stress, soil amendments to reduce salinity.

Development of resistant cultivars: This offers the cheapest, easiest to apply and most eco-friendly approach to reduce losses due to any stress.

Indirect breeding: Evaluation of materials under stress often helps in identification of a resistant line that may become an important commercial cultivar.

???????Direct breeding: It is deliberate breeding of materials for stress resistance. When a genetically variable material is grown under the stress environment, selection can be based on survival, yield and other traits contributing to stress resistance.

28.1 Introduction

The stress that is caused by biological agents or factors such as diseases, insect-pests and parasitic weeds is called biotic stress. The term disease can be broken into two words: “dis” (means negative, reverse or opposite) and “ease” (means comfort, or freedom from pain or discomfort). Hence, plant disease can be defined as any harmful alteration from the normal functioning of physiological processes. Diseases may be bacterial, fungal or viral. A wide range of insects infests various crop plants. The extent of damage caused to crop plants by various organisms is usually in the order fungi > bacteria > viruses > nematodes = insects. But the relative importance of these organisms varies to a great deal in different crops. Breeding of cultivars resistant to diseases and/or insect-pests is the cheapest, safest and most dependable means of limiting the losses due to them. Resistant cultivars form an important component of integrated management of diseases and insect-pests.

28.2 Breeding for Resistance to Biotic Stresses-Diseases

Agrios (1970) has defined disease as: “Disease is a series of invisible and visible responses of plant cells and tissues to a pathogenic microorganism or an environmental factor that result in adverse changes in form, function or integrity of plant and may lead to partial impairment or death of the plant or its parts.” The organism causing disease is called pathogen, while the plant affected by the disease is called host.

The principles underlying breeding for resistance to diseases are much the same as for other characters, except for one important difference, i.e., the diseases caused by pathogens are the product of interaction between two genetic systems, that of the host and that of the parasite. Both systems are capable of variation and evolution and affect the interaction between host and pathogen. Here the breeder is required to deal with the heritable traits of both the host plant and the pathogens invading it.

Globally, India ranked second in vegetable production and contributed 15.8 and 14% to global vegetable area and production, respectively. The production of vegetables in India is estimated to be 212.91 million tonnes in the year 2022-23 compared to 209.14 million tonnes in the year 2021-22. India ranked first in production of okra in the world (73% of world production) and second in brinjal (27.55%), cabbage (13%) and tomato (11%). The major vegetable growing states in India are West Bengal (14.1%), Uttar Pradesh (11.4%), Bihar (9.3%), Madhya Pradesh (8.0%) and Gujarat (7.1%). Vegetables are especially important crops for farmers with small holdings because an appreciably higher income per hectare can be generated by growing vegetables than from conventional staple crops (Genova et al, 2006). However, vegetables are generally considered more vulnerable than staple crops to stressful environmental conditions including extremes of temperatures, drought, salinity, alkalinity, water logging, mineral nutrient excesses and deficiency (Chinnusamy et al, 2005). These environmental stresses are likely to be exacerbated by the prevalent climatic changes in many parts of the world.
Among environmental factors, soil salinity stress is one of the main factor reducing production and productivity of vegetables in arid and semi-arid regions where irrigation water has excessive salt and annual rainfall is low. In India, about 6.73 million ha of land (~2.1% area of the country) is salt affected, out of which 2.96 million ha is saline and 3.77 million ha is sodic (Arora & Sharma 2017). About 75% of salt-affected areas exist in four states, i.e., Gujarat, Maharashtra, West Bengal, and Rajasthan (Mandal et al., 2018).

Important breeding methods can be classified as follows:
    1.    Introduction
    2.    Selection
    a.    Mass selection: Done in self pollinated crops, cross pollinated crops and dioecious crops.
    b.    Individual plant selection: Done in self pollinated crops, cross pollinated crops (line breeding) without control of pollination, artificially self pollinated lines, dioecious plants and clonally propagated lines.
    3.    Hybridization
    a.    Crossing in self pollinated crops.
    b.    Single crosses, double crosses and synthesized varieties.
    c.    Varietal crosses.

17.1 Introduction

Almond or badam is a high energy nut. Almond kernels contain lipids (52.2%), protein (20%), carbohydrate (20.4%) and water-soluble sugars (4.4%). Irrigation affects almond yield and quality. Almond quality is primarily genotype dependent. Maturity at harvest can also influence kernel quality.

12.1 Introduction

Aonla (also called Amla, Indian gooseberry) is a tropical and sub-tropical fruit. It is mainly recognized due to its high nutritive, medicinal and therapeutic properties. It contains high amount of vitamin C (500-1500 mg/100 g); it is also rich in pectin, iron, calcium and phosphorus. Dried fruits of aonla are used in curing haemorrhage, diarrhoea, chronic dysentery, diabetes, jaundice, dyspepsia, cough etc. Aonla is the main ingredient of Chavanprash and Triphala powder. The fruit contains gallic acid and leucoanthocyanin which have antioxidant property. Its cultivation is more common and widely grown in Uttar Pradesh, Gujarat, Rajasthan, Madhya Pradesh and Tamil Nadu.

14.1 Introduction

Apple is the most widely grown temperate fruit of the world. It is considered as King of temperate fruits. It is rich in carbohydrate, calcium, phosphorous, iron, potassium, vitamin B6 . Apple is rich in antioxidant, flavonoids and dietary fibre. It is recommended to reduce the incidence of dental caries. The phytonutrients and antioxidants may help reduce the risk of developing cancers, hypertensions, diabetes and heart disease. Apple is mainly used for table purpose. The various processed products are prepared from fruit, e.g., juice, jelly, preserve, canned slice, wine, concentrated juice, cider and powder. It is widely cultivated in Afghanistan, Armenia, Azerbaijan, Bhutan, China, India, Indonesia, Iraq, Iran, Israel, Japan, Jordan etc.

20.1 Introduction

Arecanut (or betel palm) is chewed as narcotic either alone or more commonly along with betel leaf. Its nut has medicinal and pharmaceutical properties. About 1/10th of world population has a habit of betel chewing. Nut contains polyphenol (11-18 %) which includes flavonols like catechin, epicatechin, leucocyanidin, fat (9-15%), polysaccharides (17-25%), fibre and protein (6- 7.5%) and some minerals like calcium, phosphorous and iron.

4.1 Introduction

Banana (Musa spp.) is one of the oldest fruits and second largest growing fruit crop in the world. It is also known as Adam’s Fig and Apple of Paradise. Important banana growing states are Maharashtra, Karnataka, Kerala, Tamil Nadu, Andhra Pradesh, Odisha, Bihar, West Bengal and Assam. In South India, banana is extensively used in all auspicious occasions such as wedding, festival and worship. Banana is a good table fruit. The cultivar Nendran is used for cooking; it is also used for preparation of halwa, sweet, chips etc. Main bud or heart of banana bunches is also used as vegetable in India, Thailand, Philippines, Malaysia and Indonesia. Musa textiles is known for strong fibre quality. Ripe banana contains about 20% sugar (but no starch) and about 4.7% protein. Further, it contains vitamins A, B, C, D and E and minerals such as K, P, Ca and Fe.

11.1 Introduction

Ber (also called Indian Jujube, Indian Plum, Chinese Apple and Masau) is an important minor fruit of India. It is most widely cultivated in Punjab, Haryana, Rajasthan, Uttar Pradesh, Madhya Pradesh, Bihar, Gujarat, Maharashtra etc. Fruits are mostly eaten fresh but other forms, such as dried, candied, squash etc. can be prepared from ber. Stem, bark, root and leaves have some medicinal values. Leaves of ber are used as fodder in dry regions.

21.1 Introduction

Cashewnut or cashew is an export-oriented crop grown for its nuts, a true drupe. It is often referred to as ‘wonder nut’. It is highly delicious and nutritive. Its nutritional composition compares well with almond, hazelnut and walnut. The economic part is kernel which is a unique combination of fats (47%), protein (21%), carbohydrates, minerals and vitamins. Nutritionally it is equivalent to milk, egg and meat, it is good appetizer, an excellent nerve tonic, stimulant, body builder. India is the largest producer, processor and exporter; and second largest consumer in the world. Cashew is grown in Kerala, Karnataka, Goa and Maharashtra along the west coast and in Tamil Nadu, Andhra Pradesh, Odisha and West Bengal along the east coast.

5.1 Introduction

Citrus is one of the most important fruit crops in the world. It is grown in the tropical and sub-tropical regions of the world. The top ten citrus producing countries are China, India, Nigeria, Brazil, Mexico, USA, Spain, Egypt, Italy and Argentina. The citrus industry in India is the third fruit industry after mango and banana. The important groups of crops in citrus are sweet orange, mandarins, limes, lemons, grape fruits and pummelo. The most important commercial Citrus species grown in India are mandarin (Citrus reticulata Blanco.), followed by sweet orange (Citrus sinensis (L.) Osbeck) and acid lime (Citrus aurantifolia Swingle). The major citrus producing states are Maharashtra, Karnataka, Andhra Pradesh and Punjab.

19.1 Introduction

Coconut is valued most for its nut. The other components are coconut water, kernel, shell and husk. Coconut water is one of the most refreshing drinks. It is also known for its medicinal value. It increases blood circulation in kidneys, causes profuse diuresis and eliminates mineral poisoning. It is a good substitute for saline glucose under gastroenteritis conditions. The coconut has been regarded as ‘Kalpavriksha’ because all the parts of the palm is useful to mankind in one way or other. The dried kernels commonly called as copra is the richest source of vegetable fat (60-67 % oil). Coconut is major source of edible oil. India is the largest producer of coir and its product. The major coconut growing areas of the world are found in Asia, Oceania, West Indies, Central South America, East and West Africa. In India, coconut is commercially grown in Kerala, Karnataka, Tamil Nadu, Andhra Pradesh, Gujarat, Odisha and West Bengal.

23.1 Introduction

Coffee is the second important popular beverage for its aroma and mild stimulant action. Its dried beans are roasted, ground and brewed to make a stimulating and refreshing beverage. Use of coffee has evolved from original chewing of leaves and beans of the plant to relieve pain, hunger and fatigue to the present more sophisticated uses, like espresso and decaffeinated coffees. Its use was first discovered in Arabia around mid-15th century.

6.1 Introduction

Grapes are being grown commercially in the tropics. The region between the tropics of Cancer and Capricorn which was once considered to be unsuitable for growing quality grapes is producing good quality grapes. Tropical countries where grapes are grown are India, Indonesia, the Philippines, Thailand, Taiwan, South China, Australia, Brazil, Columbia, Ecuador, Mexico, Venezuela, West Indies, Kenya, Nigeria and Zimbabwe. In India, about 90% of grapes are being produced in the tropical states including Maharashtra, Andhra Pradesh, Karnataka and Tamil Nadu. There are three main commercial uses of grape, viz., table, raisin and wine.

8.1 Introduction

Guava is the fourth important fruit crop next to mango, banana and citrus. It is known as the ‘Apple of tropics’. It is very rich and cheap source of Vitamin C and also contains a fair amount of calcium. Important guava growing states are Uttar Pradesh, Bihar, Madhya Pradesh and Maharashtra. Allahabad district of Uttar Pradesh has the reputation of growing the best quality of guava fruits in the world. The importance of guava is due to the fact that it is the hardy fruit which can be grown in alkaline and poorly drained soil. Guava fruits are used for making jam, jelly and various culinary purposes. The fruit can be canned in sugar syrup.

———————— is a thornless and seedless selection from Kagzi lime.

A. Pramalini            B. Vikram

C. Chakradhar        D. Jai devi

10.1 Introduction

Litchi is a delicious fruit and generally consumed as a table fruit. It is one of the most popular fruits in India. It is also recognized as “Queen of Fruits”. It is a subtropical evergreen fruit crop. Its climatic requirements are highly specific. Thus, its commercial cultivation is restricted to only a few subtropical countries of the world. The major litchi producing countries are China, India, Taiwan, Thailand and Vietnam. In India, the important litchi growing states are Bihar, Jharkhand, West Bengal, Tripura, Uttar Pradesh, Uttarakhand, Chhattisgarh, Punjab and Himachal Pradesh. Litchi makes an excellent canned fruit and highly flavoured squash is prepared from inferior fruits.

3.1 Introduction

The Mango (Mangifera indica L.) belonging to the family Anacardiaceae in order Sapindales, is amongst the most important tropical fruits of the world. In India, it is also known as the ‘king of fruits’ owing to the delicious quality of the fruit rich in vitamins and minerals. Its long period of domestication is evident from its mention in the ancient Indian scriptures. In India, mango has been valued not only for religious consideration but also for its importance in the economic and cultural life of the society. Mango is rich in Vitamin A and C, flavonoids, carotenes, glucosides, sterols, terpenes, aromatic acids, essential oils, fatty acids and phenolics. It is a powerful nutritive fruit, containing most of the essential substances needed by the human body. The most important single factor limiting the production of mango fruit at the present time is the erratic fruiting habits of this species. Some cultivars produce a good crop in most years and a poor crop in other years, while still other cultivars produce only a few fruits each year. For many years new cultivars have been selected and grown on the basis of size, appearance, and dessert quality with very little attention given to their production capabilities. The improvement of mango needs to explore new recombinants primarily by means of exploiting the breeding methodologies. Diversity or heterogeneity is the main character desired for breeding either natural or manmade. It is required to have vast genetic pool to get new combinations of desired nature and developing new hybrids. It has been mentioned earlier that the development of mango in India is the result of selection from the amateur gardeners. Now, the scientists have developed certain hybrids of mango.

24.1 Introduction

In India, the commercial cultivation of oil palm started during 1970. Today about 16 countries of tropics cultivate oil palm. The palm oil has been safe and nutritious source of edible oil for human consumption for 1000 years. Palm oil and its liquid fraction palmolein is consumed worldwide as cooking oil. Malaysia, Indonesia, Nigeria and Columbia started to grow oil palm in a large scale during 1980. The commercial producing countries are Liberia, Peru, Angola, Senegal, Ghana, etc.

7.1 Introduction

Papaya is a fruit crop grown in kitchen garden, backyards of home and orchards. It is grown as a filler crop in orchard. India is the largest producer of papaya in the world. It is also cultivated in Brazil, Mexico, Australia, Hawaii, Malaysia, Taiwan, Peru, Florida, Gold Coast, South Africa and Bangladesh. In India, it is widely cultivated in Uttar Pradesh, Bihar, Karnataka, Jharkhand and Madhya Pradesh. The ripe fruits of papaya are consumed throughout the tropics and sub-tropics. Fruits are also used in the preparation of jam, soft drinks, ice cream flavour, crystallized fruits and syrups. Unripe fruits are commonly used as vegetables. Papain is prepared from the latex of immature fruits. It is a proteolytic enzyme used for tendering of meat, preparation of chewing gum and cosmetics. The ripe fresh fruit is rich source of vitamin A, carbohydrates and minerals. The fruits are prescribed for piles, dyspepsia, liver, spleen and digestive disorders. Roots are used as nerve tonic. The plants flower five months after planting and fruits are ready five months after flowering. So, it has attracted attention of fruit growers to cultivate on a large scale in the country.

16.1 Introduction

Peach is one of the important fruits grown in warm temperate zones of the world. Fruits are rich in protein, sugar, minerals and vitamins. Peaches are highly valued as table fruit for their attractive colour and palatability. Canned peaches, dried frozen preserves, jams, nectar are principal processed products of peach. It is grown mainly in Jammu and Kashmir, Himachal Pradesh, Punjab, Uttarakhand, Nilgiri hills, Jharkhand and North-Eastern States.

15.1 Introduction

Pear is one of the important fruit crops that belong to the family Rosaceae. In the world pear is cultivated in China, USA, Argentina, Italy, Turkey, Spain, India, South Africa, Japan, Belgium etc. It is grown under a wide variety of climatic regimes ranging from cold dry temperate hilly conditions to warm humid sub-tropical conditions on the plains of northern India. In India pear is widely cultivated in Punjab, Himachal Pradesh, Jammu and Kashmir, Uttar Pradesh, Uttarakhand etc.

9.1 Introduction

Pineapple is one of the most important commercial fruits of the world. It is good source of vitamin A, B and fairly rich in vitamin C, minerals like calcium, phosphorous and iron. Fresh fruit is used as dessert. Fruit is also used for juice, squash, jam, mixed jam and in canned form. Fruit core is used for preparing candy. Some cultivars of pineapple are used for extraction of fibre. The important pineapple growing states are Assam, West Bengal, Tripura, Kerala, Andhra Pradesh, Bihar, Karnataka and Madhya Pradesh.

13.1 Introduction

Pomegranate (Punica granatum L.), an economically important fruit plant species, belongs to the family Punicaceae. It is an important fruit crop of arid and semiarid regions of the world and has been believed to be the symbol of fertility and abundance. Pomegranate is a highly remunerative crop for replacing subsistence farming and alleviating poverty. The plant is drought tolerant, winter hardy and can thrive well under desert condition. It is a good source of protein, carbohydrate, minerals, antioxidants, vitamins A, B and C, and has also been used in controlling diarrhoea, hyperacidity, tuberculosis, leprosy, abdominal pain and fever. Pomegranate juice contains antioxidants such as soluble polyphenols, tannins, anthocyanins and can be used in the treatment of cancer and chronic inflammation. It has enjoyed a reputation for its healthy dietetic and medicinal properties.

22.1 Introduction

Tea is the most important non-alcoholic beverage gaining further popularity as an important “health drink”. It is consumed as a morning drink by 2/3rd of world population. Tea was initially used as a medicine and subsequently as beverage and now has proven to be a potential raw material for pharmaceutical industry. Tea is mainly consumed in the form of ‘fermented tea’ or ‘black tea’, ‘non-fermented tea’ or ‘green tea’ and ‘semi-fermented tea’ or ‘oolong tea’. Green leaves are also used as vegetables such as ‘leppet tea’ in Burma and ‘meing tea’ in Thailand. Tea leaves have more than 700 chemical constituents among which flavonoids, amino acids, vitamins (C, E, K), caffeine and polysaccharides are important to human health. The stimulating effect of tea is due to caffeine (1.25-4.5%). Per cup value of caffeine in tea is three and half times less than that of coffee and is less harmful than coffee. Importantly, the vitamin C content in leaves is comparable to that of lemon.

18.1 Introduction

Among nut crops, ‘Walnut’ or ‘Akhrot’ is the most important crop. It has immense commercial importance even in the areas situated in remotes without having good market and transport infrastructure. Because of long storability, growers have opportunity to hold the produce at room temperature for two to three months easily and sell out at intended time. Walnut is rich in protein, fats and vitamins, especially vitamin B6 . The walnut kernel contains about 90% PUFA of which 26% is the Omega-3 fatty acid, alpha-linolenic acid. But the problem with this type of kernel is that higher the percentage of PUFA in it, the more is the chances of getting the oil oxidized and thereby reducing the shelf life of the walnut kernel.

2.1 Introduction

The different breeding strategies (methods) that can be employed for improvement of fruit crops include introduction, open pollinated selection, clonal selection, hybrid breeding, polyploidy breeding, mutation breeding, molecular breeding, in vitro breeding and transgenic breeding. Combining traditional approaches with biotechnological approaches will speed up the rate of genetic improvement. Thus, what is required is to adopt a coordinated approach involving breeders, biotechnologists, horticulturists, pathologists, entomologists and statistician for making the cultivation of fruit crops more scientific and more profitable and thereby contributing more to improving the health of the people in a sustainable manner.